Conjunctio
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Conjunctio
''Conjunctio'' is an extinct genus of dissorophid temnospondyl amphibian from the early Permian of New Mexico. The type species, ''Conjunctio multidens'', was named by paleontologist Robert L. Carroll in 1964. History of study The holotype specimen was found in 1911 in the Lower Permian Abo Formation in New Mexico ) , population_demonym = New Mexican ( es, Neomexicano, Neomejicano, Nuevo Mexicano) , seat = Santa Fe , LargestCity = Albuquerque , LargestMetro = Tiguex , OfficialLang = None , Languages = English, Spanish ( New Mexican), Navajo, Ker .... It consists of the skull and postcranial material including femora, humeri, scapulae, pelvis, a section of the vertebral column, and osteoderms. It was originally described by Case et al. (1913) as a referred specimen of ''Aspidosaurus'' (sometimes "''Broiliellus''") ''novomexicanus'' but was subsequently determined to be a distinct species by Carroll (1964). Carroll also identified a third specimen of ''Conjunctio'', ...
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Dissorophids
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be ...
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Dissorophid
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be d ...
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Dissorophidae
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be d ...
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Scapanops
''Scapanops'' is an extinct genus of dissorophid temnospondyl amphibian known from the Early Permian Nocona Formation of north-central Texas, United States. It contains only the type species ''Scapanops neglecta'', which was named by Rainer R. Schoch and Hans-Dieter Sues in 2013. ''Scapanops'' differs from other dissorophids in having a very small skull table, which means that its eye sockets are unusually close to the back of the skull. The eye sockets are also very large and spaced far apart. ''Scapanops'' was probably small-bodied (around long) with a proportionally large head and short trunk and tail. Like other dissorophids, it probably spent most of its life on land. Description The only known skull of ''Scapanops'' is long. ''Scapanops'' is distinguished from other dissorophids by five unique features or autapomorphies: a large spacing between the orbits or eye sockets, giving the skull an oval-shaped outline; a very small skull table behind the orbits; a long snout t ...
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Dissorophus
''Dissorophus'' (DI-soh-ROH-fus) (meaning "double roof" for two layers of armor) is an extinct genus of temnospondyl amphibian that lived during the Early Permian Period about 273 million years ago. Its fossils have been found in Texas and in Oklahoma in North America. Its heavy armor and robust build indicate ''Dissorophus'' was active on land, similar to other members of the clade Dissorophidae that are known from the Late Carboniferous to the Early Permian periods. ''Dissorphus'' is distinguished by its small body size, disproportionately large head and short trunk. The American paleontologist Edward Drinker Cope first briefly described ''Dissorophus'' in 1895,Cope. E.D. 1895. A batrachian armadillo. ''American Naturalist'' 29:99/ref> likely deriving the genus name from Ancient Greek δισσός/dissos "double" and ὀροφή/orophe "roof" to refer to the double layer of armor formed by horizontal "spinous branches" at the top of the neural spines of the vertebrae that "tou ...
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Broiliellus Texensis
''Broiliellus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. ''Broiliellus'' is most closely related to the genus '' Dissorophus'', and both have been placed in the subfamily Dissorophinae. ''Broiliellus'' is known from five species from the Early Permian: the type species is ''Broiliellus texensis,'' and the other species are ''Broiliellus brevis,'' ''Broiliellus olsoni, Broiliellus arroyoensis,'' and ''Broiliellus reiszi''. An additional species, ''Broiliellus novomexicanus'', which was originally named '' Aspidosaurus novomexicanus'', is now thought to fall outside the genus as a member of the subfamily Eucacopinae. History of study ''Broiliellus'' was first named by American paleontologist S.W. Williston in 1914 based on two nearly complete skulls in articulation with postcranial material from the early Permian of Texas; this species was given the name ''Broiliellus texensis'', the genus name being for the German paleontologist Ferdin ...
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Aspidosaurus
''Aspidosaurus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. Description Like other dissorophids, ''Aspidosaurus'' species had a single row of plates formed by expansions of the neural spines.A Description of ''Aspidosaurus novomexicanus'' Williston
Permo-Carboniferous Vertebrates from New Mexico, p7-11. Retrieved 2011-09-11.


Taxonomy

In 1911, Paul Miller discovered the remains of various dissorophid bones in that were attributed to a new species, ''Aspidosaurus novomexicanus''. The skull closely resembled that of a specimen of ''

Broiliellus Brevis
''Broiliellus'' is an extinct genus of dissorophoid temnospondyl within the family Dissorophidae. ''Broiliellus'' is most closely related to the genus ''Dissorophus'', and both have been placed in the subfamily Dissorophinae. ''Broiliellus'' is known from five species from the Early Permian: the type species is ''Broiliellus texensis,'' and the other species are ''Broiliellus brevis,'' ''Broiliellus olsoni, Broiliellus arroyoensis,'' and ''Broiliellus reiszi''. An additional species, ''Broiliellus novomexicanus'', which was originally named ''Aspidosaurus novomexicanus'', is now thought to fall outside the genus as a member of the subfamily Eucacopinae. History of study ''Broiliellus'' was first named by American paleontologist S.W. Williston in 1914 based on two nearly complete skulls in articulation with postcranial material from the early Permian of Texas; this species was given the name ''Broiliellus texensis'', the genus name being for the German paleontologist Ferdinand B ...
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Dissorophinae
Dissorophinae is a subfamily of dissorophid temnospondyl Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphi ...s that includes '' Dissorophus'' and '' Broiliellus''. Phylogeny Below is the cladogram from Schoch (2012): References * Permian temnospondyls {{temnospondyli-stub ...
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Eucacopinae
Eucacopinae is an extinct clade (evolutionary grouping) of dissorophid temnospondyls. Eucacopines differ from the other main group of dissorophids, the Dissorophinae, in having more lightly built skeletons and more knobby skulls. The subfamily was originally named Cacopinae, but since the name was already established for a group of living microhylid frogs in 1931, the name was changed to Eucacopinae in 2013. Eucacopinae is a stem-based taxon defined as the most inclusive clade containing the species '' Cacops apsidephorus'' but not '' Dissorophus multicinctus'', which belongs to Dissorophinae. According to the most recent phylogenetic analyses of Dissorophidae, Eucacopinae includes the basal ("primitive") species ''Conjunctio multidens'' and '' Scapanops neglecta'' from the southwestern United States and a more derived ("advanced") group including several species of ''Cacops'' (also from the southwestern United States) and the Russian genera ''Kamacops'' and '' Zygosaurus''. Der ...
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Cacops Woehri
''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids) whose ontogeny is known. ''Cacops'' fossils were almost exclusively known from the Cacops Bone Bed of the Lower Permian Arroyo Formation of Texas for much of the 20th century. New material collected from the Dolese Brothers Quarry, near Richards Spur, Oklahoma in the past few decades has been recovered, painting a clearer picture of what the animal looked and acted like. History of discovery ''Cacops aspidephorus'' is the most famous dissorophid, in part due to a majority of its skeleton having been known for over a century. Over 50 specimens have been found in the ''Cacops'' Bone Bed in Baylor County, Texas, which is now flooded by the dammed Lake Kemp. However, many of the specimens are covered in calcite, which penetrates the bone tis ...
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Kamacops
''Kamacops'' is a genus of dissorophid temnospondyls known from the Middle to Late Permian of Russia that was described by Yuri Gubin in 1980. It is known from a single species, ''Kamacops acervalis,'' material of which is currently reposited in the Paleontological Institute of the Russian Academy of Sciences. ''Kamacops'' is one of the youngest dissorophids, along with '' Iratusaurus'' and '' Zygosaurus'' from Russia and ''Anakamacops'' from China and was one of the largest known dissorophids, with an estimated skull length of 24–30 cm. It is typically recovered as being most closely related to ''Zygosaurus'' and to the North American ''Cacops ''Cacops'' ("ugly look" for its strange appearance), is a genus of dissorophid temnospondyls from the Kungurian stage of the early Permian of the United States. ''Cacops'' is one of the few olsoniforms (dissorophids and the larger trematopids) ...''''.'' A detailed study of the braincase region was performed by Schoch (1999), ...
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