Ciliate Biology
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Ciliate Biology
The ciliates are a group of alveolates characterized by the presence of hair-like organelles called cilia, which are identical in structure to eukaryotic flagella, but are in general shorter and present in much larger numbers, with a different undulating pattern than flagella. Cilia occur in all members of the group (although the peculiar Suctoria only have them for part of their life cycle) and are variously used in swimming, crawling, attachment, feeding, and sensation. Ciliates are an important group of protists, common almost anywhere there is water—in lakes, ponds, oceans, rivers, and soils. About 4,500 unique free-living species have been described, and the potential number of extant species is estimated at 27,000–40,000. Included in this number are many ectosymbiotic and endosymbiotic species, as well as some obligate and opportunistic parasites. Ciliate species range in size from as little as 10 µm in some colpodeans to as much as 4 mm in length in some gel ...
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Ediacaran
The Ediacaran Period ( ) is a geological period that spans 96 million years from the end of the Cryogenian Period 635 million years ago (Mya), to the beginning of the Cambrian Period 538.8 Mya. It marks the end of the Proterozoic Eon, and the beginning of the Phanerozoic Eon. It is named after the Ediacara Hills of South Australia. The Ediacaran Period's status as an official geological period was ratified in 2004 by the International Union of Geological Sciences (IUGS), making it the first new geological period declared in 120 years. Although the period takes its name from the Ediacara Hills where geologist Reg Sprigg first discovered fossils of the eponymous Ediacaran biota in 1946, the type section is located in the bed of the Enorama Creek within Brachina Gorge in the Flinders Ranges of South Australia, at . The Ediacaran marks the first appearance of widespread multicellular fauna following the end of Snowball Earth glaciation events, the so-called Ediacaran biota, ...
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Prostomatea
Prostomatea is a class of ciliates. It includes the genera ''Coleps ''Coleps'' is a genus of ciliates in the class Prostomatea with barrel-shaped bodies surrounded by regularly arranged plates composed of calcium carbonate. Description Species of ''Coleps'' can grow up to 250 µm in length, but are usua ...'' and '' Pelagothrix''. References Further reading Intramacronucleata Ciliate classes {{ciliate-stub ...
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Biological Life Cycle
In biology, a biological life cycle (or just life cycle or lifecycle when the biological context is clear) is a series of changes in form that an organism undergoes, returning to the starting state. "The concept is closely related to those of the life history, development and ontogeny, but differs from them in stressing renewal." Transitions of form may involve growth, asexual reproduction, or sexual reproduction. In some organisms, different "generations" of the species succeed each other during the life cycle. For plants and many algae, there are two multicellular stages, and the life cycle is referred to as alternation of generations. The term life history is often used, particularly for organisms such as the red algae which have three multicellular stages (or more), rather than two.Dixon, P.S. 1973. ''Biology of the Rhodophyta.'' Oliver & Boyd. Life cycles that include sexual reproduction involve alternating haploid (''n'') and diploid (2''n'') stages, i.e., a change of pl ...
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Suctoria
Suctoria are ciliates that become sessile in their developed stage and then lose their redundant cilia. They feed by extracellular digestion. They were originally thought to feed by suction – hence their name. In fact, they use specialized microtubules to ensnare and manipulate their prey. They live in both freshwater and marine environments, including some that live on the surface of aquatic animals, and typically feed on other ciliates. Instead of a single cytostome, each cell feeds by means of several specialized tentacles. These are supported by microtubules and phyllae, and have toxic extrusomes called haptocysts at the tip, which they attach to prey. They then suck the prey's cytoplasm directly into a food vacuole inside the cell, where they digest and absorb its contents. Most suctoria are around 15-30 μm in size, with a non-contractile stalk and often a lorica or shell. Suctoria reproduce primarily by budding, producing swarmers that lack both tentacles and ...
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Flagellum
A flagellum (; ) is a hairlike appendage that protrudes from certain plant and animal sperm cells, and from a wide range of microorganisms to provide motility. Many protists with flagella are termed as flagellates. A microorganism may have from one to many flagella. A gram-negative bacterium ''Helicobacter pylori'' for example uses its multiple flagella to propel itself through the mucus lining to reach the stomach epithelium, where it may cause a gastric ulcer to develop. In some bacteria the flagellum can also function as a sensory organelle, being sensitive to wetness outside the cell. Across the three domains of Bacteria, Archaea, and Eukaryota the flagellum has a different structure, protein composition, and mechanism of propulsion but shares the same function of providing motility. The Latin word means " whip" to describe its lash-like swimming motion. The flagellum in archaea is called the archaellum to note its difference from the bacterial flagellum. Eukaryotic ...
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Cilia
The cilium, plural cilia (), is a membrane-bound organelle found on most types of eukaryotic cell, and certain microorganisms known as ciliates. Cilia are absent in bacteria and archaea. The cilium has the shape of a slender threadlike projection that extends from the surface of the much larger cell body. Eukaryotic flagella found on sperm cells and many protozoans have a similar structure to motile cilia that enables swimming through liquids; they are longer than cilia and have a different undulating motion. There are two major classes of cilia: ''motile'' and ''non-motile'' cilia, each with a subtype, giving four types in all. A cell will typically have one primary cilium or many motile cilia. The structure of the cilium core called the axoneme determines the cilium class. Most motile cilia have a central pair of single microtubules surrounded by nine pairs of double microtubules called a 9+2 axoneme. Most non-motile cilia have a 9+0 axoneme that lacks the central pair of mi ...
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Alveolate
The alveolates (meaning "pitted like a honeycomb") are a group of protists, considered a major clade and Biological classification, superphylum within Eukarya. They are currently grouped with the stramenopiles and Rhizaria among the protists with tubulocristate mitochondria, the group being referred to as SAR supergroup, SAR. Characteristics The most notable shared characteristic is the presence of cortical (near the surface) alveoli (sacs). These are flattened vesicle (biology), vesicles (sacs) arranged as a layer just under the cell membrane, membrane and supporting it, typically contributing to a flexible pellicle (thin skin). In armored dinoflagellates they may contain stiff plates. Alveolates have mitochondrion, mitochondria with tubular cristae (invaginations), and cells often have pore-like intrusions through the cell surface. The group contains free-living and parasitic organisms, predatory flagellates, and photosynthetic organisms. Almost all sequenced mitochondrial g ...
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Parablepharismea
Parablepharismea is a class of free-living marine and brackish anaerobic ciliates that form a major clade of obligate anaerobes within the SAL group ( Spirotrichea, Armophorea, and Litostomatea), together with the classes Muranotrichea and Armophorea Armophorea is a class of ciliates in the subphylum Intramacronucleata. . It was first resolved in 2004 and comprises three orders: Metopida, Clevelandellida, and Armophorida. Previously members of this class were thought to be heterotrichs beca .... Parablepharismea are medium to large, elongated ciliates with navicular outline and holotrichous somatic ciliature composed of dikinetids without postciliodesmata. Their oral ciliature is composed of bipartite paroral membrane and adoral zone of membranelles. They host a thick coat of prokaryotic ectosymbionts and cytoplasmic endosymbionts. They are found in micro-oxic marine to brackish habitats. Parablepharismea may be related to the deep sea Cariacotrichea ciliates. This class ...
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Muranotrichea
Muranotrichea is a class of free-living marine anaerobic ciliates, that, together with the classes Parablepharismea and Armophorea, form a major clade of obligate anaerobes within the SAL group ( Spirotrichea, Armophorea, and Litostomatea). Muranotrichea are medium to large, elongated holotrichous ciliates with contractile body and somatic ciliature composed of dikinetids. Their oral ciliature includes an elongated key-hole shaped peristome, an adoral zone of membranelles spiraling rightward around distinctly neck-like anterior body part; and heteromorphic paroral membrane . They inhabit micro-oxic marine to brackish sediments and host prokaryotic ectosymbionts and endosymbionts. The class contains a single order Muranotrichida and family Muranotrichidae. Muranotrichidae include two species, ''Muranothrix'' ''gubernata'' and ''Thigmothrix strigosa''. Muranotrichea together with Parablepharismea Parablepharismea is a class of free-living marine and brackish anaerobic ciliat ...
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Spirotrich
The spirotrichs are a large and diverse group of ciliate protozoa. They typically have prominent oral cilia in the form of a series of polykinetids, called the adoral zone of membranelles, beginning anterior to the oral cavity and running down to the left side of the mouth. There may also be one or two paroral membranes on its right side. The body cilia are fused to form polykinetids called cirri in some, and are sparse to absent in others. Forms with cirri are common throughout soil, freshwater, and marine environments. Individuals tend to be flattened, with cirri confined to the ventral surface. These are variously used for crawling over objects, acting as feet, swimming, or assisting in food capture. They are generally divided into hypotrichs and stichotrichs, but were originally all considered hypotrichs. Forms with sparse or absent body cilia tend to be smaller and are mostly marine, but a few are common in freshwater. Again, they are generally divided into oligotrich ...
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