Cearadactylus
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Cearadactylus
''Cearadactylus'' is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of ''Cearadactylus'' dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is ''C. atrox'', described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek ''daktylos'', "finger", a reference to the wing finger of pterosaurs. The Latin ''atrox'' means "frightful", a reference to the fearsome dentition of the species. Discovery The holotype of ''Cearadactylus'' is MN 7019-V (formerly CB-PV-F-O93), which was discovered in the Romualdo Formation of the Santana Group in the Araripe plateau of northeastern Brazil. It consists of a single skull with a length of .Leonardi, G. & Borgomanero, G. (1985). "''Cearadactylus atrox'' nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Arar ...
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Cearadactylus Model
''Cearadactylus'' is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of ''Cearadactylus'' dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is ''C. atrox'', described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek ''daktylos'', "finger", a reference to the wing finger of pterosaurs. The Latin ''atrox'' means "frightful", a reference to the fearsome dentition of the species. Discovery The holotype of ''Cearadactylus'' is MN 7019-V (formerly CB-PV-F-O93), which was discovered in the Romualdo Formation of the Santana Group in the Araripe plateau of northeastern Brazil. It consists of a single skull with a length of .Leonardi, G. & Borgomanero, G. (1985). "''Cearadactylus atrox'' nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Ara ...
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Cearadactylus
''Cearadactylus'' is a genus of large anhanguerid pterodactyloid pterosaur from the Romualdo Formation of Brazil, South America. Fossil remains of ''Cearadactylus'' dated back to the Albian stage of the Early Cretaceous period, about 112 million years ago. The only known species is ''C. atrox'', described and named in 1985 by Giuseppe Leonardi and Guido Borgomanero. The name refers to the Brazilian state Ceará, and combines this with Greek ''daktylos'', "finger", a reference to the wing finger of pterosaurs. The Latin ''atrox'' means "frightful", a reference to the fearsome dentition of the species. Discovery The holotype of ''Cearadactylus'' is MN 7019-V (formerly CB-PV-F-O93), which was discovered in the Romualdo Formation of the Santana Group in the Araripe plateau of northeastern Brazil. It consists of a single skull with a length of .Leonardi, G. & Borgomanero, G. (1985). "''Cearadactylus atrox'' nov. gen., nov. sp.: novo Pterosauria (Pterodactyloidea) da Chapada do Arar ...
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Coloborhynchus
''Coloborhynchus'' is a genus of pterodactyloid pterosaur belonging to the family Anhangueridae, though it has also been recovered as a member of the Ornithocheiridae in some studies. ''Coloborhynchus'' is known from the Lower Cretaceous of England (Valanginian age, 140 to 136 million years ago), and depending on which species are included, possibly the Albian and Cenomanian ages (113 to 93.9 million years ago) as well. ''Coloborhynchus'' was once thought to be the largest known toothed pterosaur, however, a specimen of the closely related '' Tropeognathus'' is now thought to have had a larger wingspan. History and classification Like many ornithocheiroid pterosaurs named during the 19th century, ''Coloborhynchus'' has a highly convoluted history of classification. Over the years numerous species have been assigned to it, and often, species have been shuffled between ''Coloborhynchus'' and related genera by various researchers. In 1874 Richard Owen, rejecting the creation by ...
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Romualdo Formation
The Romualdo Formation is a geologic Konservat-Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, previously designated as the Romualdo Member of the Santana Formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau. It was discovered by Johann Baptist von Spix in 1819. The strata were deposited during the Aptian stage of the Early Cretaceous in a lacustrine rift basin with shallow marine incursions of the proto-Atlantic. At that time, the South Atlantic was opening up in a long narrow shallow sea. The Romualdo Formation earns the designation of Lagerstätte due to an exceedingly well preserved and diverse fossil faunal assemblage. Some 25 species of fossil fishes are often found with stomach contents preserved, enabling paleontologists to study predator–prey relationships in this ecosystem. There are also fine examples of pterosaurs, reptiles a ...
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Anhanguera (pterosaur)
''Anhanguera'' () is a genus of pterodactyloid pterosaur known from the Early Cretaceous (Albian age, 125 to 112 million years ago) Romualdo Formation of Brazil and the Late Cretaceous ( Cenomanian age, 98 to 92.5 million years ago) Kem Kem Group of Morocco. This pterosaur is closely related to ''Ornithocheirus'', but belongs in the family Anhangueridae.Campos, D. de A., and Kellner, A. W. (1985). "Um novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formação Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. The generic name comes from the Tupi words ''añanga'', meaning "spirit protector of the animals" + ''wera'' "bygone". Description ''Anhanguera'' was a fish-eating animal with a wingspan of about .Aureliano, T., Ghilardi, A. M., Duque, R. R., & Barreto, A. M. (2014). ON THE OCCURRENCE OF PTEROSAURIA IN EXU, PERNAMBUCO (LOWER CRETACEOUS ROMUALDO FORMATION, ARARIPE BASI ...
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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested '' Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct cla ...
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Anhanguerid
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested '' Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct cl ...
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Giancarlo Ligabue
Giancarlo Ligabue (30 October 1931 – 25 January 2015) was an Italian paleontologist, scholar, politician and businessman. Born in Venice, Ligabue graduated in Economics at the Ca' Foscari University of Venice and in Geology at la Sorbonne, and later he also received five honorary degrees from the universities of Bologna, Venice, Modena, Lima and Ashgabat. He participated or directed over 130 expeditions around the world, and he made several paleontological discoveries, such as the ''Ligabueino'', a noasaurid dinosaur named after him, and such as the unearthed deposits of hominid and dinosaurs fossils in the Ténéré desert. He collaborated with Piero Angela to several science documentaries. He was president of the Natural History Museum of Venice and founder of the Ligabue Study and Research Centre in Venice. Ligabue was also businessman in the field of supplies and services for ships. He was the president of the basketball team Reyer Venezia Mestre between the early sixtie ...
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Teeth
A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, tearing food, for defensive purposes, to intimidate other animals often including their own, or to carry prey or their young. The roots of teeth are covered by gums. Teeth are not made of bone, but rather of multiple tissues of varying density and hardness that originate from the embryonic germ layer, the ectoderm. The general structure of teeth is similar across the vertebrates, although there is considerable variation in their form and position. The teeth of mammals have deep roots, and this pattern is also found in some fish, and in crocodilians. In most teleost fish, however, the teeth are attached to the outer surface of the bone, while in lizards they are attached to the inner surface of the jaw by one side. In cartilaginous fi ...
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Piscivore
A piscivore () is a carnivorous animal that eats primarily fish. The name ''piscivore'' is derived . Piscivore is equivalent to the Greek-derived word ichthyophage, both of which mean "fish eater". Fish were the diet of early tetrapod evolution (via water-bound amphibians during the Devonian period); insectivory came next; then in time, the more terrestrially adapted reptiles and synapsids evolved herbivory. Almost all predatory fishes (most sharks, tuna, billfishes, pikes etc.) are obligated piscivores. Some non-piscine aquatic animals, such as whales, sea lion and crocodilians, are not completely piscivorous, often also preying on invertebrates, marine mammals, waterbirds and even wading land animals in addition to fish; while others, such as the bulldog bat and gharial, are strictly dependent on fish for food. Some creatures, including cnidarians, octopuses, squid, spiders, cetaceans, grizzly bears, jaguars, wolves, snakes, turtles and sea gulls, may have fish as significa ...
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Fabio Marco Dalla Vecchia
Fabio is a given name descended from Latin ''Fabius'' and very popular in Italy and Latin America (due to Italian migration). Its English equivalent is Fabian. The name is written without an accent in Italian and Spanish, but is usually accented in Portuguese as ''Fábio'' (with the diminutive Fabinho or Fabiano). The presence or absence of the written accent does not affect pronunciation. First name A–K * Fabio (DJ), drum-and-bass DJ and producer from the UK * Fabio Armiliato (born 1956), Italian operatic tenor * Fábio Aurélio (born 1979), Brazilian footballer * Fábio Bahia (Fábio Júnior Nascimento Santana, born 1983), currently playing for Goiás * Fabio Bencivenga, Italian water polo player * Jud "Fabio" Birza, winner of ''Survivor: Nicaragua'' * Fabio Borini, Italian footballer * Fábio Camilo de Brito (Nenê, born 1975), currently playing for Coritiba Foot Ball Club * Fabio Cannavaro, former captain of the Italy national team * Fabio Capello, Italian manager of the ...
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Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0.9 Ma (million years ago). The Albian is preceded by the Aptian and followed by the Cenomanian. Stratigraphic definitions The Albian Stage was first proposed in 1842 by Alcide d'Orbigny. It was named after Alba, the Latin name for River Aube in France. A Global Boundary Stratotype Section and Point (GSSP), ratified by the IUGS in 2016, defines the base of the Albian as the first occurrence of the planktonic foraminiferan '' Microhedbergella renilaevis'' at the Col de Pré-Guittard section, Arnayon, Drôme, France. The top of the Albian Stage (the base of the Cenomanian Stage and Upper Cretaceous Series) is defined as the place where the foram species '' Rotalipora globotruncanoides'' first appears in the stratigraphic column. The Alb ...
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