Carcharodini
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Carcharodini
The Carcharodini are a tribe in the skipper butterfly subfamily Pyrginae. They are a very diverse but quite plesiomorphic and inconspicuous group distributed throughout the tropics.Brower (2009) Formerly, when only four tribes of Pyrginae were recognized, they were included in the Pyrgini, which at that time contained a massive number of genera. But the Pyrginae have since been reorganized to make them and their tribes monophyletic, leading most modern authors to treat the Carcharodini as distinct tribe. However, the old circumscription of the Pyrgini was by and large just as correct from a phylogenetic perspective. The tribe of the Pyrgini ''sensu lato'' most closely related to the Pyrgini ''sensu stricto'' are the strikingly different Achlyodidini. As many consider it desirable to treat this lineage as a distinct tribe, the Carcharodini naturally need to be considered distinct too. Genera These genera belong to the tribe Carcharodini: * '' Agyllia'' Grishin, 2020 * '' Arteu ...
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Iliana
''Iliana'' is a genus of skippers in the family Hesperiidae. It was described in 1937 by Ernest Layton Bell, with ''Iliana romulus'' as type species. The genus has a neotropical distribution. Distribution Species mainly occur in Peru, Brazil and Bolivia, with one species, ''Iliana remus'', additionally known from French Guiana. Taxonomy Placement within the Hesperiidae The placement of ''Iliana'' within the Hesperiidae is not fully resolved. It has traditionally been placed within the Pyrgini, but Warren et al. (2008, 2009) place it in Carcharodini, which has been followed by many but not all subsequent publications including the genus. Mielke and Casagrande (2016) argue for its placement to remain in tribe Pyrgini until more in-depth reclassification has taken place. Species At the time the genus was erected, it contained two species, ''Iliana remus'' and ''Iliana romulus'', with the latter assigned as type species. Both species were at the time newly described by Bell from ...
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Hesperopsis
''Hesperopsis'' is a genus of butterflies in the skipper family, Hesperiidae. Species include:''Hesperopsis''.
Integrated Taxonomic Information System (ITIS) *'' Hesperopsis alpheus'' (Edwards, 1876) *'' Hesperopsis gracielae'' (MacNeill, 1970) *''
Hesperopsis libya ''Hesperopsis libya'', the Mojave sootywing, Mohave sootywing, Great Basin sootywing or Lena sooty wing, is a butterfly of the family Hesperiidae. It is found ...
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Gorgopas (butterfly)
''Gorgopas'' is a genus of skippers in the family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... ReferencesNatural History Museum Lepidoptera genus database Carcharodini Hesperiidae genera Taxa named by Osbert Salvin Taxa named by Frederick DuCane Godman {{Pyrginae-stub ...
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Gomalia
''Gomalia'' is a genus of marbled skippers in the butterfly family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... There are three described species in ''Gomalia''. ''Gomalia albofasciata'' was formerly considered subspecies of ''Gomalia elma'', and ''Gomalia jeanneli'' was formerly considered a synonym of ''Gomalia elma''. They were both reinstated as full species as a result of genomic research published in 2020. Species These species belong to the genus ''Gomalia'': * '' Gomalia albofasciata'' Moore, 1879 (Asian marbled skipper) * '' Gomalia elma'' (Trimen, 1862) (green-marbled skipper or African marbled skipper) * '' Gomalia jeanneli'' (Picard, 1949) References Further reading * Carcharodini Taxa named by Frederic Moore Hesperiidae genera {{pyr ...
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Agyllia
''Agyllia'' is a genus of spread-wing skippers in the butterfly family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy ..., erected in 2020. There are at least three described species in ''Agyllia'', all found in Africa. These species were formerly members of the genus ''Spialia''. Species These species belong to the genus ''Agyllia'': * '' Agyllia agylla'' (Trimen, 1889) * '' Agyllia asterodia'' (Trimen, 1864) * '' Agyllia kituina'' (Karsch, 1896) References Further reading * Carcharodini {{pyrginae-stub ...
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Bolla (butterfly)
''Bolla'' is a genus of spread-winged skippers found from Mexico to South America.''Bolla''
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Species

Listed alphabetically.Jeffrey Glassberg (2007) A Swift Guide to the Butterflies of Mexico and Central America. Sunstreak Books Inc. . pp.183-185. * '' Bolla antha'' Evans, 1953 * '' Bolla atahuallpai'' (Lindsey, 1925) – Atahuallpa bolla * '' Bolla boliviensis'' (Bell, 1937) – Bolivian bolla * ...
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Ernsta
''Ernsta'' is a genus of spread-wing skippers in the butterfly family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... There are about 14 described species in ''Ernsta'', found mainly in Africa. These species were formerly members of the genus ''Spialia''. Species These species belong to the genus ''Ernsta'': * '' Ernsta bifida'' (Higgins, 1924) * '' Ernsta colotes'' (Druce, 1875) (Bushveld Sandman) * '' Ernsta confusa'' (Higgins, 1924) (Confusing Sandman) * '' Ernsta delagoae'' (Trimen, 1898) (Delagoa Sandman) * '' Ernsta depauperata'' (Strand, 1911) (Wandering Sandman) * '' Ernsta dromus'' (Plötz, 1884) (Forest Sandman) * '' Ernsta mangana'' (Rebel, 1899) (Arabian Grizzled Skipper) * '' Ernsta nanus'' (Trimen, 1889) (Dwarf Sandman) * '' Ernsta paula'' (Higgins, 19 ...
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Cyclosemia
''Cyclosemia'' is a genus of skippers in the family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... ReferencesNatural History Museum Lepidoptera genus database Carcharodini Hesperiidae genera Taxa named by Paul Mabille {{Pyrginae-stub ...
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Conognathus
''Conognathus'' is a genus of skippers in the family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... ReferencesNatural History Museum Lepidoptera genus database Carcharodini Hesperiidae genera Taxa named by Baron Cajetan von Felder Taxa named by Rudolf Felder {{Pyrginae-stub ...
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Arteurotia
''Arteurotia'' is a genus of skipper butterflies genus in the family Hesperiidae Skippers are a family of the Lepidoptera (moths and butterflies) named the Hesperiidae. Being diurnal, they are generally called butterflies. They were previously placed in a separate superfamily, Hesperioidea; however, the most recent taxonomy .... Species Two species are recognised in the genus ''Arteurotia'': * '' Arteurotia celendris'' Hewitson, 1878 * '' Arteurotia tractipennis'' Butler & Druce, 1872 - starred skipper - Mexico, Costa Rica and Panama References * Carcharodini Hesperiidae genera Monotypic butterfly genera {{Pyrginae-stub ...
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Achlyodidini
The Achlyodidini (sometimes – but apparently in errorWahlberg ''fide'' Savela (2008) – spelled Achylodidini) are a tribe in the skipper butterfly subfamily Pyrginae. Many species of them are notable for their (by skipper butterfly standards) gaudy colors and the unusually-shaped wings, which look as if the forewing tips have been squarely cut off. They are only found in the Americas, with few occurring out of tropical areas.Brower & Warren (2009) Formerly, when only four tribes of Pyrginae were recognized, they were included in the Pyrgini, which at that time contained a massive number of genera. But the Pyrginae have since been reorganized to make them and their tribes monophyletic, leading most modern authors to treat the Achlyodidini as distinct tribe. However, they are the tribe most closely related to the Pyrgini '' sensu stricto'', and the old circumscription of the Pyrgini was by and large just as correct from a phylogenetic perspective. The other two trib ...
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Carcharodus
''Carcharodus'' is a Palearctic genus of skippers in the family Hesperiidae. In genomic research published in 2020, it was determined that six of the eight species in ''Carcharodus'' were more closely related to those in the genus ''Muschampia'', and were transferred to that genus. This left only two species in ''Carcharodus''. Description This genus includes medium-sized, brown spotted skippers. Distribution and habitat They are widespread in central and southern Europe, North Africa, and in temperate Asia from northern India to southern Siberia. These skippers can be found in meadows, trails and other sunny, warm locations. They fly in the sunshine and often visit flowers to drink nectar. The larvae feed on various herbs of several different plant families, including Malvaceae Malvaceae, or the mallows, is a family of flowering plants estimated to contain 244 genera with 4225 known species. Well-known members of economic importance include okra, cotton, cacao and duria ...
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