Sawflies
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Sawflies
Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyletic, consisting of several basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the thorax. Some sawflies are Batesian mimics of wasps and bees, and the ovipositor can be mistaken for a stinger. Sawflies vary in le ...
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Sawfly (Nematus Miliaris) Larvae
Sawflies are the insects of the suborder Symphyta within the order Hymenoptera, alongside ants, bees, and wasps. The common name comes from the saw-like appearance of the ovipositor, which the females use to cut into the plants where they lay their eggs. The name is associated especially with the Tenthredinoidea, by far the largest superfamily in the suborder, with about 7,000 known species; in the entire suborder, there are 8,000 described species in more than 800 genera. Symphyta is paraphyletic, consisting of several basal groups within the order Hymenoptera, each one rooted inside the previous group, ending with the Apocrita which are not sawflies. The primary distinction between sawflies and the Apocrita – the ants, bees, and wasps – is that the adults lack a "wasp waist", and instead have a broad connection between the abdomen and the thorax. Some sawflies are Batesian mimics of wasps and bees, and the ovipositor can be mistaken for a stinger. Sawflies vary in len ...
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Hymenoptera
Hymenoptera is a large order of insects, comprising the sawflies, wasps, bees, and ants. Over 150,000 living species of Hymenoptera have been described, in addition to over 2,000 extinct ones. Many of the species are parasitic. Females typically have a special ovipositor for inserting eggs into hosts or places that are otherwise inaccessible. This ovipositor is often modified into a stinger. The young develop through holometabolism (complete metamorphosis)—that is, they have a wormlike larval stage and an inactive pupal stage before they mature. Etymology The name Hymenoptera refers to the wings of the insects, but the original derivation is ambiguous. All references agree that the derivation involves the Ancient Greek πτερόν (''pteron'') for wing. The Ancient Greek ὑμήν (''hymen'') for membrane provides a plausible etymology for the term because species in this order have membranous wings. However, a key characteristic of this order is that the hindwings are co ...
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Xyeloidea
The Xyelidae are a comparatively species-poor family of sawflies comprising about 80 extant species in five genera worldwide and is the only family in the superfamily Xyeloidea. The fossil record of the family is extensive, comprising more than 120 species and including the oldest fossil Hymenoptera species dating back to the Triassic, between 245 and 208 million years ago. Xyelidae are to be regarded as living fossils since they represent one of the oldest lineages of insects and include still extant forms. The extant species occur in the Northern Hemisphere, especially in boreal regions of the Holarctic, though there are a few Oriental species. Two genera and about 15 species occur in Europe.Taeger, A., Blank, S.M. & Liston, A. D. 2006: European Sawflies (Hymenoptera: Symphyta) – A Species Checklist for the Countries. Pp. 399-504. In: Blank, S.M., Schmidt, S. & Taeger, A. (eds): Recent Sawfly Research: Synthesis and Prospects. Goecke & Evers, Keltern.Blank, S.M. 2002: Biosys ...
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Orussoidea
Orussoidea is a superfamily of sawflies. It contains the living family Orussidae, as well as the extinct families Burmorussidae and Paroryssidae. They are the group of sawflies closest to the Apocrita, the group containing wasps, bees and ants, with both groups together forming the clade Euhymenoptera. Like most members of Apocrita, but unlike other sawflies, members of the superfamily are parasitoids. Taxonomy * Burmorussidae Qi Zhang, Dmitry S. Kopylov and A. P. Rasnitsyn, 2020 Burmese amber, Myanmar, Late Cretaceous ( Cenomanian) ** '' Burmorussus'' Qi Zhang, Dmitry S. Kopylov and A. P. Rasnitsyn, 2020 ** '' Cretorussus'' Jouault, Perrichot & Nel, 2021 * Paroryssidae Martynov 1925 ** '' Microryssus'' Rasnitsyn 1968 Karabastau Formation Kazakhstan, Middle-Late Jurassic (Callovian/ Oxfordian) ** '' Paroryssus'' Martynov 1925 Karabastau Formation Kazakhstan, Middle-Late Jurassic (Callovian/ Oxfordian) ** '' Praeoryssus'' Rasnitsyn 1968 Karabastau Formation Kazakhstan, M ...
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Pamphilioidea
The Pamphilioidea are a small superfamily within the Symphyta (the sawflies), containing some 250 living species restricted to the temperate regions of Eurasia and North America. These hymenopterans share the distinctive feature of a very large, almost prognathous head, which is widest ventrally. The superfamily contains two extant families. The Pamphiliidae are the leaf-rolling or web-spinning sawflies such as ''Acantholyda'', ''Neurotoma'', and '' Pamphilius'' whose larvae eat plants such as conifers; the adults have simple filiform antennae. The Megalodontesidae include genera such as ''Megalodontes'' and several fossil groups. Their larvae eat herbaceous plants, while the adults have serrate or pectinate antennae. References Bibliography * , in Zhang, Z.-Q. (ed.) Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013) * Rasnitsyn, Alexandr P.; Zhang, Haichun & Wang, Bo (2006): Bizarre fossil insects: web-spinnin ...
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Tenthredo Mesomela
''Tenthredo mesomela'' is a sawfly species of the family Tenthredinidae (common sawflies), subfamily Tenthredininae. Description ''Tenthredo mesomela'' can reach a length of about . These sawflies have a yellow to apple green body with black head, thorax and upperside of the abdomen, while pronotum and scutellum are yellow. They are distinguished from the very similar species of the genus '' Rhogogaster'' by the position of the eyes and by their black pterostigma ('' Rhogogaster'' species have a green stigma). The adults can be encountered from May through July feeding on small insects and on nectar and pollen of flowers (especially on Apiaceae species). The larvae feed at night on leaves of buttercup (''Ranunculus'' species) and '' Persicaria'' species (''Polygonaceae The Polygonaceae are a family of flowering plants known informally as the knotweed family or smartweed—buckwheat family in the United States. The name is based on the genus ''Polygonum'', and was first ...
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Tenthredinoidea
The Tenthredinoidea are the dominant superfamily of sawflies within the Symphyta, containing some 8,400 species worldwide, primarily in the family Tenthredinidae. All known larvae are phytophagous, and a number are considered pests. The included extant families share the distinctive features of a medially narrowed pronotum, paired protibial spurs, and the loss of the transverse mesonotal groove. The superfamily also includes two extinct families. Meicai and Haiyan (1998) identified 66 extant tribes and 17 subfamilies. Taxonomy Families * Argidae Konow, 1890 (58 genera, 897 spp.) * Blasticotomidae Thomson, 1871 (3 genera, 13 spp.) * Cimbicidae W. Kirby, 1837 (16 genera, 182 spp.) * Diprionidae Rohwer, 1910 (11 genera, 136 spp.) *Pergidae The Pergidae are a moderate-sized family of sawflies occurring in the Western Hemisphere and the Australasian Region. The Pergidae are, with almost 450 described species, the third-largest family of Symphyta after the Tenthredin ...
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Paraphyly
In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic group (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of Synapomorphy and apomorphy, synapomorphies and symplesiomorphy, symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term was coined by Willi Hennig to apply to well-known taxa like Reptilia (reptiles) which, as commonly named and traditionally defined, is paraphyletic with respect to mammals and birds. Reptilia contains the last common ancestor of reptiles a ...
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Siricoidea
The superfamily Siricoidea is an archaic group of the order Hymenoptera, consisting of six families (four extinct) of xylophagous sawflies. The group is well represented in early Tertiary and Mesozoic The Mesozoic Era ( ), also called the Age of Reptiles, the Age of Conifers, and colloquially as the Age of the Dinosaurs is the second-to-last era of Earth's geological history, lasting from about , comprising the Triassic, Jurassic and Creta ... times, but a number of living taxa remain, including the family Anaxyelidae, which has recently been linked to this group (it was previously placed in the Xyeloidea). The female ovipositor is typically long and projects posteriorly, and is used to drill into wood. Families These six families belong to the superfamily Siricoidea: * Siricidae (horntails) * Xiphydriidae (wood wasps, sometimes treated as a separate superfamily, Xiphydrioidea) * † Daohugoidae * † Protosiricidae * † Pseudosiricidae * † Sinosiricidae Refe ...
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Cephoidea
Cephoidea is a small superfamily within the Symphyta, commonly referred to as stem sawflies, containing some 100 species in 10 genera in the living family, Cephidae, plus another 17 genera in the extinct family Sepulcidae. Most species occur in the Northern Hemisphere, especially in Eurasia. The larvae are stem borers in various plants, especially grasses, but sometimes other herbaceous plants, shrubs, or trees. A few are pests of cereal grains (e.g. '' Cephus cinctus'', which attacks wheat). They are exceptionally slender for symphytans, often resembling other types of wasps, and they are the only Symphyta which lack cenchri. They are sometimes postulated to be the sister taxon to the Apocrita, though the Orussidae The Orussidae or the parasitic wood wasps represent a small family of sawflies ("Symphyta"). Currently, about 93 extant and four fossil species are known. They take a key position in phylogenetic analyses of Hymenoptera, because they form the si ... are more commo ...
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Proleg
A proleg is a small, fleshy, stub structure found on the ventral surface of the abdomen of most larval forms of insects of the order Lepidoptera, though they can also be found on other larval insects such as sawflies and a few other types of insects. In all the orders in which they appear, mainly Hymenoptera and Lepidoptera, prolegs of any form evolved independently of each other by convergent evolution. Prolegs of lepidopteran larvae have a small circle of gripping hooks, called "crochets". The arrangement of the crochets can be helpful in identification to family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ... level. Although the point has been debated, prolegs are not widely regarded as true legs, derived from the primitive uniramous limbs. Certainly in their morpholog ...
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Karatavitoidea
Karatavitidae is an extinct family of sawflies, known from the Jurassic period, they are the only members of the superfamily Karatavitoidea. While once proposed to be grouped with the Orussoidea in the infraorder Orussomorpha, they are now considered to be the closest relatives of clade containing Orussoidea and Apocrita. There are about 7 genera in Karatavitidae. Genera These seven genera belong to the family Karatavitidae: * † '' Grimmaratavites'' Rasnitsyn, Ansorge & Zhang, 2006 Green Series, Germany, Early Jurassic (Toarcian) * † '' Karatavites'' Rasnitsyn, 1963 Karabastau Formation Kazakhstan, Middle-Late Jurassic (Callovian/ Oxfordian), Daohugou The Haifanggou Formation (), also known as the Jiulongshan Formation (), is a fossil-bearing rock deposit located near Daohugou () village of Ningcheng County, in Inner Mongolia, northeastern China. The formation consists of coarse conglomerate ..., China, Callovian * † '' Postxiphydria'' Rasnitsyn & Zhang, 2010 Karabas ...
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