Reptiliomorphs
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Reptiliomorphs
Reptiliomorpha (meaning reptile-shaped; in PhyloCode known as ''Pan-Amniota'') is a clade containing the amniotes and those tetrapods that share a more recent common ancestor with amniotes than with living amphibians (lissamphibians). It was defined by Michel Laurin (2001) and Vallin and Laurin (2004) as the largest clade that includes ''Homo sapiens'', but not '' Ascaphus truei'' (tailed frog). Laurin and Reisz (2020) defined Pan-Amniota as the largest total clade containing ''Homo sapiens'', but not ''Pipa pipa'', '' Caecilia tentaculata'', and ''Siren lacertina''. The informal variant of the name, "reptiliomorphs", is also occasionally used to refer to stem-amniotes, i.e. a grade of reptile-like tetrapods that are more closely related to amniotes than they are to lissamphibians, but are not amniotes themselves; the name is used in this meaning e.g. by Ruta, Coates and Quicke (2003). An alternative name, "Anthracosauria", is also commonly used for the group, but is confusingly ...
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Tetrapod
Tetrapods (; ) are four-limbed vertebrate animals constituting the superclass Tetrapoda (). It includes extant and extinct amphibians, sauropsids ( reptiles, including dinosaurs and therefore birds) and synapsids (pelycosaurs, extinct therapsids and all extant mammals). Tetrapods evolved from a clade of primitive semiaquatic animals known as the Tetrapodomorpha which, in turn, evolved from ancient lobe-finned fish (sarcopterygians) around 390 million years ago in the Middle Devonian period; their forms were transitional between lobe-finned fishes and true four-limbed tetrapods. Limbed vertebrates (tetrapods in the broad sense of the word) are first known from Middle Devonian trackways, and body fossils became common near the end of the Late Devonian but these were all aquatic. The first crown-tetrapods (last common ancestors of extant tetrapods capable of terrestrial locomotion) appeared by the very early Carboniferous, 350 million years ago. The specific aquatic ancestors ...
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Solenodonsaurus
''Solenodonsaurus'' ("single-tooth lizard") is an extinct genus of reptiliomorphs that lived in what is now Czech Republic, during the Westphalian stage. Description ''Solenondosaurus'' had snout-vent length with a skull length . ''Solenodonsaurus'' shows a curious mix of characters making it difficult to place phylogenetically. The teeth lack labyrinthodont folding of the enamel, and it skull has a much smaller otic notch than seen in other reptiliomorph amphibians. Yet general build ties it in with the Diadectomorpha. Laurin, M. and Rize R.R. (1999): A new study of ''Solenodonsaurus janenschi'', and a reconsideration of amniote origins and stegocephalian evolution. ''Canadian Journal of Earth Science'', no 36 (8): pp 1239–1255 (1999)/ref> Paleobiology ''Solenodonsaurus'' was likely best adapted to life on land, as opposed to living in an aquatic environment like many other early tetrapods. The limbs and pelvis are incomplete in all known specimens of ''Solenodonsaurus' ...
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Embolomeri
Embolomeri is an order of tetrapods or stem-tetrapods, possibly members of Reptiliomorpha. Embolomeres first evolved in the Early Carboniferous ( Mississippian) Period and were the largest and most successful predatory tetrapods of the Late Carboniferous ( Pennsylvanian) Period. They were specialized semiaquatic predators with long bodies for eel-like undulatory swimming. Embolomeres are characterized by their vertebral centra, which are formed by two cylindrical segments, the pleurocentrum at the rear and intercentrum at the front. These segments are equal in size. Most other tetrapods have pleurocentra and intercentra which are drastically different in size and shape. Embolomeres were among the earliest large carnivorous tetrapods, with members such as the crocodilian-like ''Proterogyrinus'' appearing in the Visean stage of the Carboniferous. They declined in diversity during the Permian period, though at least one representative ('' Archeria)'' was common in the Early Permian ...
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Gephyrostegidae
Gephyrostegidae is an extinct family of reptiliomorph tetrapods from the Late Carboniferous including the genera '' Gephyrostegus'', '' Bruktererpeton'', and '' Eusauropleura''. ''Gephyrostegus'' is from the Czech Republic, ''Brukterepeton'' is from Germany, and ''Eusauropleura'' is from the eastern United States The United States of America (U.S.A. or USA), commonly known as the United States (U.S. or US) or America, is a country primarily located in North America. It consists of 50 states, a federal district, five major unincorporated territorie .... References Reptiliomorphs Pennsylvanian first appearances Pennsylvanian extinctions {{carboniferous-animal-stub ...
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Mississippian (geology)
The Mississippian ( , also known as Lower Carboniferous or Early Carboniferous) is a subperiod in the geologic timescale or a subsystem of the geologic record. It is the earlier of two subperiods of the Carboniferous period lasting from roughly 358.9 to 323.2 million years ago. As with most other geochronologic units, the rock beds that define the Mississippian are well identified, but the exact start and end dates are uncertain by a few million years. The Mississippian is so named because rocks with this age are exposed in the Mississippi Valley. The Mississippian was a period of marine transgression in the Northern Hemisphere: the sea level was so high that only the Fennoscandian Shield and the Laurentian Shield were dry land. The cratons were surrounded by extensive delta systems and lagoons, and carbonate sedimentation on the surrounding continental platforms, covered by shallow seas. In North America, where the interval consists primarily of marine limestones, it is treate ...
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Casineria
''Casineria'' is an extinct genus of tetrapod which lived about 340-334 million years ago in the Mississippian epoch of the Carboniferous period. Its generic name, ''Casineria'', is a latinization of Cheese Bay, the site near Edinburgh, Scotland where the holotype fossil was found. When originally described in 1999, it was identified as a transitional fossil noted for its mix of basal (amphibian-like) and advanced (reptile-like) characteristics, putting it at or very near the origin of the amniotes, the group containing all mammals, birds, modern reptiles, and other descendants of their reptile-like common ancestor. However, the sole known fossil is lacking key elements such as a skull, making exact analysis difficult. As a result, the classification of ''Casineria'' has been more controversial in analyses conducted since 1999. Other proposed affinities include a placement among the lepospondyls, seymouriamorphs, " gephyrostegids", or as a synonym of '' Caerorhachis'', another ...
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Michel Laurin
Michel Laurin is a Canadian-born French vertebrate paleontologist whose specialities include the emergence of a land-based lifestyle among vertebrates, the evolution of body size and the origin and phylogeny of lissamphibians. He has also made important contributions to the literature on phylogenetic nomenclature. As an undergraduate, he worked in the laboratory of Robert L. Carroll and earned his Ph.D. at the University of Toronto under the direction of Robert R. Reisz; his thesis concerned the osteology of seymouriamorphs. His 1991 review of diapsid phylogeny provided the broadest review of the subject up to that date. In 1995, Laurin and Reisz coauthored a widely cited article providing evidence that the synapsids are the sister group of all other amniotes. He later worked on untangling the phylogeny of the Stegocephalia, a group with a notoriously difficult phylogeny. He later moved to France; since 1998, he has been a CNRS researcher at the Muséum National d'Histoire Naturel ...
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Lissamphibia
The Lissamphibia is a group of tetrapods that includes all modern amphibians. Lissamphibians consist of three living groups: the Salientia (frogs, toads, and their extinct relatives), the Caudata (salamanders, newts, and their extinct relatives), and the Gymnophiona (the limbless caecilians and their extinct relatives). A fourth group, the Allocaudata, was moderately successful, spanning 160 million years from the Middle Jurassic to the Early Pleistocene, but became extinct two million years ago. For several decades, this name has been used for a group that includes all living amphibians, but excludes all the main groups of Paleozoic tetrapods, such as Temnospondyli, Lepospondyli, Embolomeri, and Seymouriamorpha. Some scientists have concluded that all of the primary groups of modern amphibians—frogs, salamanders and caecilians—are closely related. Some writers have argued that the early Permian dissorophoid ''Gerobatrachus hottoni'' is a lissamphibian. If it is not, the earl ...
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Amphibian
Amphibians are tetrapod, four-limbed and ectothermic vertebrates of the Class (biology), class Amphibia. All living amphibians belong to the group Lissamphibia. They inhabit a wide variety of habitats, with most species living within terrestrial animal, terrestrial, fossorial, arboreal or freshwater aquatic ecosystems. Thus amphibians typically start out as larvae living in water, but some species have developed behavioural adaptations to bypass this. The young generally undergo metamorphosis from larva with gills to an adult air-breathing form with lungs. Amphibians use their skin as a secondary respiratory surface and some small terrestrial salamanders and frogs lack lungs and rely entirely on their skin. They are superficially similar to reptiles like lizards but, along with mammals and birds, reptiles are amniotes and do not require water bodies in which to breed. With their complex reproductive needs and permeable skins, amphibians are often ecological indicators; in re ...
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Amniotes
Amniotes are a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds, and extinct parareptiles and non-avian dinosaurs) and synapsids (including pelycosaurs and therapsids such as mammals). They are distinguished from the other tetrapod clade — the amphibians — by the development of three extraembryonic membranes ( amnion for embryoic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage), thicker and more keratinized skin, and costal respiration (breathing by expanding/constricting the rib cage). All three main features listed above, namely the presence of an amniotic buffer, water-impermeable cutes and a robust respiratory system, are very important for amniotes to live on land as true terrestrial animals – the ability to reproduce in locations away from water bodies, better homeostasis in drier environments, and more efficient air respiration to power terrestrial locomotions, although they migh ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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PhyloCode
The ''International Code of Phylogenetic Nomenclature'', known as the ''PhyloCode'' for short, is a formal set of rules governing phylogenetic nomenclature. Its current version is specifically designed to regulate the naming of clades, leaving the governance of species names up to the rank-based nomenclature codes ('' ICN'', ''ICNCP'', '' ICNP'', '' ICZN'', '' ICVCN''). The ''PhyloCode'' is associated with the International Society for Phylogenetic Nomenclature (ISPN). The companion volume, ''Phylonyms'', establishes 300 taxon names under ''PhyloCode'', serving as examples for those unfamiliar with the code. RegNum is an associated online database for registered clade names. The ''PhyloCode'' regulates phylogenetic nomenclature by providing rules for deciding which associations of names and definitions are considered established, which of those will be considered homonyms or synonyms, and which one of a set of synonyms or homonyms will be considered accepted (generally the one regi ...
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