Paracrocodylomorphs
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Paracrocodylomorphs
Paracrocodylomorpha is a clade of pseudosuchian archosaurs. The clade includes the diverse and unusual group Poposauroidea as well as the generally carnivorous and quadrupedal members of Loricata, including modern crocodylians. Paracrocodylomorpha was named by paleontologist J. Michael Parrish in 1993, although the group is now considered to encompass more reptiles than his original definition intended. The most recent definition of Paracrocodylomorpha, as defined by Sterling Nesbitt in 2011, is "the least inclusive clade containing ''Poposaurus'' and ''Crocodylus niloticus'' (the Nile crocodile). Most groups of paracrocodylomorphs became extinct at the end of the Triassic period, with the exception of the crocodylomorphs, from which crocodylians such as crocodiles and alligators evolved in the latter part of the Mesozoic. History and definition Parrish (1993) defined Paracrocodylomorpha as the last common ancestor of "Poposauridae" and Crocodylomorpha and all of its descendants ...
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Ticinosuchus BW
''Ticinosuchus'' is an extinct genus of suchian archosaur from the Middle Triassic (Anisian - Ladinian) of Switzerland and Italy. Description One of only a handful of fossil reptiles that have been found in Switzerland, ''Ticinosuchus'' (meaning "Ticino crocodile" due to its origin from the Swiss canton Ticino) was about long, and its whole body, even the belly, was covered in thick, armoured scutes. These scutes were sometimes considered to have been staggered, alternating between several rows. However, some studies refute this claim, instead purporting that the scutes were aligned in neat rows, with a one-to-one assignment of scutes to vertebrae. The structure of the hips shows that its legs were placed under the body almost vertically. Coupled with the development of a calcaneus and a specialized ankle joint, this would have made ''Ticinosuchus'' a fast runner, unlike most earlier reptiles. ''Ticinosuchus'' is thought to be very close to or possible even the same species t ...
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Vytshegdosuchus
''Vytshegdosuchus'' is an extinct genus of paracrocodylomorph archosaur known from the Early Triassic (latest Olenekian stage) Yarenskian Gorizont of the Komi Republic of the European section of Russia. It contains a single species, ''Vytshegdosuchus zheshartensis''. ''Vytshegdosuchus'' was named by Andreii Sennikov in 1988. Discovery ''Vytshegdosuchus'' is known from the holotype PIN 3361/134, a right ilium, and possibly from the paratype PIN 3361/127, a partial femur. These specimens were collected from the a fossil assemblage called the ''Parotosuchus'' fauna of the Upper Yarenskian Gorizont, in the Zheshart locality along the Vychegda River in Aikino district of the Komi Republic. A pterygoid and postcranial material from the Gam locality, a cervical vertebra (PIN 3369/139) from another locality and postcrania from the Mezhog locality were also assigned to ''Vytshegdosuchus'' by Sennikov (1999). Nesbitt (2009) argued that the specimens referred to ''Vytshegd ...
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Mambawakale
''Mambawakale'' is a genus of large sized basal paracrocodylomorph from the Manda Beds of Tanzania. It was informally named ''Pallisteria'' before being officially published under its current name almost 60 years after its discovery. It contains a single species, ''Mambawakale ruhuhu''. History and naming In 1963, following the independence of Tanzania, Alan Charig participated in a joint expedition of the Natural History Museum, London and the University of London, as well as researchers from Uganda, South Africa and Edinburgh, to Tanzania and Zambia. The expedition heavily relied on the support of locals who had discovered the localities and fossils within them, although they went unnamed in the field notes. The fossils discovered in 1963 were collected and stored in the Natural History Museum of London. Among these fossils was the incomplete skull of a large crocodylomorph, noted for its large size and informally referred to as ''Pallisteria angustimentum'' (after Charig's f ...
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Loricata
Loricata is a clade of archosaur reptiles that includes crocodilians and some of their Triassic relatives, such as ''Postosuchus'' and ''Prestosuchus''. More specifically, Loricata includes Crocodylomorpha (the persistent archosaur subset which crocodilians belong to) and most "rauisuchians", a paraphyletic grade of large terrestrial pseudosuchians which were alive in the Triassic period and ancestral to crocodylomorphs. Loricata is one branch of the group Paracrocodylomorpha; the other branch is the clade Poposauroidea, an unusual collection of strange "rauisuchians" including bipedal, herbivorous, and sail-backed forms. The vast majority of typical "rauisuchians", which were usually quadrupedal predators, occupy basal (early-branching) rungs of Loricata leading up to crocodylomorphs. History and Classification Loricata was initially named in a completely different context by German naturalist Blasius Merrem in his 1820 ''Versuch eines Systems der Amphibien''. Merrem consid ...
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Saurosuchus
''Saurosuchus'' (meaning "lizard crocodile") is an extinct genus of large loricatan pseudosuchian archosaur that lived in South America during the Late Triassic period. It was a heavy, ground-dwelling, quadrupedal carnivore, likely being the apex predator in the Ischigualasto Formation. Discovery and naming The holotype, PVL 206, was discovered by Galileo J. Scaglia and Leocadio Soria in 1957, lying in a greenish sandstone on the Cancha de Bochas Member of the Ischigualasto Formation in the Ischigualasto-Villa Unión Basin in northwestern Argentina. It consists of a nearly complete, but deformed skull. ''Saurosuchus'' was formally described and named later in 1959 by Osvaldo. A. Reig. The generic name, ''Saurosuchus'', is derived from the Greek (, meaning lizard) and (, meaning crocodile). The specific name, ''galilei'', is in honour to Galileo J. Scaglia, who unearthed and prepared the holotype. Referred specimens ''Saurosuchus'' is known from numerous specimens coming fr ...
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Batrachotomus
''Batrachotomus'' is a genus of prehistoric archosaur. Fossils of this animal have been found in southern Germany and dated from the Ladinian stage of the Middle Triassic period, around 242 to 237 million years ago. ''Batrachotomus'' was described by palaeontologist David J. Gower 22 years after its discovery. The locality where ''Batrachotomus'' lived was a swampy region and the name comes from the Greek ''batrachos/βάτραχος'' (frog) and ''tome/τομή'' (cutting, slicing), which refers to its preying on the large amphibian ''Mastodonsaurus''.Gower (1999), p. 6. In contrast with sprawling reptiles, like crocodiles, this large carnivore was very agile with locomotor superiority due to its erect stance. A remarkable feature seen on its back was a row of paired, flattened bony plates. ''Batrachotomus'' was possibly an early relative of ''Postosuchus'',Gower (1999), p. 1. which lived during the dawn of the dinosaurs. Description ''Batrachotomus'' was a heavily built, ...
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Early Triassic
The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which is a unit in chronostratigraphy. The Early Triassic is the oldest epoch of the Mesozoic Era. It is preceded by the Lopingian Epoch (late Permian, Paleozoic Era) and followed by the Middle Triassic Epoch. The Early Triassic is divided into the Induan and Olenekian ages. The Induan is subdivided into the Griesbachian and Dienerian subages and the Olenekian is subdivided into the Smithian and Spathian subages. The Lower Triassic series is coeval with the Scythian Stage, which is today not included in the official timescales but can be found in older literature. In Europe, most of the Lower Triassic is composed of Buntsandstein, a lithostratigraphic unit of continental red beds. The Early Triassic and partly also the Middle Triassic span the in ...
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Rauisuchia
"Rauisuchia" is a paraphyletic group of mostly large and carnivorous Triassic archosaurs. Rauisuchians are a category of archosaurs within a larger group called Pseudosuchia, which encompasses all archosaurs more closely related to crocodilia Crocodilia (or Crocodylia, both ) is an order of mostly large, predatory, semiaquatic reptiles, known as crocodilians. They first appeared 95 million years ago in the Late Cretaceous period ( Cenomanian stage) and are the closest living ...ns than to birds and other dinosaurs. First named in the 1940s, Rauisuchia was a name exclusive to Triassic archosaurs which were generally large (often ), carnivorous, and quadrupedal with a pillar-erect hip posture, though exceptions exist for all of these traits. Rauisuchians, as a traditional Taxonomy, taxonomic group, were considered distinct from other Triassic archosaur groups such as early dinosaurs, Phytosaur, phytosaurs (crocodile-like carnivores), Aetosaur, aetosaurs (armored herbivor ...
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Last Common Ancestor
In biology and genetic genealogy, the most recent common ancestor (MRCA), also known as the last common ancestor (LCA) or concestor, of a set of organisms is the most recent individual from which all the organisms of the set are descended. The term is also used in reference to the ancestry of groups of genes (haplotypes) rather than organisms. The MRCA of a set of individuals can sometimes be determined by referring to an established pedigree. However, in general, it is impossible to identify the exact MRCA of a large set of individuals, but an estimate of the time at which the MRCA lived can often be given. Such ''time to most recent common ancestor'' (''TMRCA'') estimates can be given based on DNA test results and established mutation rates as practiced in genetic genealogy, or by reference to a non-genetic, mathematical model or computer simulation. In organisms using sexual reproduction, the ''matrilineal MRCA'' and ''patrilineal MRCA'' are the MRCAs of a given population c ...
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Poposauridae
Poposauridae is a family of large carnivorous archosaurs which lived alongside dinosaurs during the Late Triassic. They were around long. Poposaurids are known from fossil remains from North and South America. While originally believed to be theropod dinosaurs (they mirrored the theropods in a number of respects, such as features of the skull and bipedal locomotion), cladistic analysis has shown them to be more closely related to crocodiles. An early cladistic analysis of crocodylotarsan archosaurs included ''Poposaurus'', ''Postosuchus'', ''Teratosaurus'', and ''Bromsgroveia'' within Poposauridae.Parrish JM. 1993. Phylogeny of the Crocodylotarsi, with reference to archosaurian and crurotarsan monophyly. ''Journal of Vertebrate Paleontology'' 13: 287-308. However, later studies found ''Teratosaurus'' to be a rauisuchid.Long RA, Murry PA. 1995. Late Triassic (Carnian and Norian) tetrapods from the Southwestern United States. ''New Mexico Museum of Natural History & Science Bul ...
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Crocodylomorpha
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, crocodylomorphs were far more diverse than they are now. Triassic forms were small, lightly built, active terrestrial animals. The earliest and most primitive crocodylomorphs are represented by " sphenosuchians", a paraphyletic assemblage containing small-bodied forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, Crocodylomorphs morphologically diversified into numerous niches, including into the aquatic and marine realms. Evolutionary history When their extinct species and stem group are examined, the crocodylian lineage (clade Pseudosuchia, formerly Crurotarsi) proves to have been a very diverse and adaptive ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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