Cystopteris
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Cystopteris
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Laurentiana
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Utahensis
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Tennesseensis
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Reevesiana
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Douglasii
''Cystopteris'' is a genus of ferns in the family Cystopteridaceae. These are known generally as bladderferns or fragile ferns. They grow in temperate areas worldwide. This is a very diverse genus and within a species individuals can look quite different, especially in harsh environments where stress stunts their growth. They hybridize easily with each other and identifying an individual can be challenging. In general these are rhizomatous perennials which grow in rocks or soil. Their leaves are multiply pinnate, in that each leaflet is divided into smaller parts. The sori are usually rounded and covered in an inflated bladder-like indusium. Species Species include: *''Cystopteris alpina'' *''Cystopteris bulbifera'' - bulblet fern *'' Cystopteris chinensis'' (Ching) R.Wei & X.C.Zhang (syn. ''Cystoathyrium chinense'' Ching) *'' Cystopteris diaphana'' *'' Cystopteris dickieana'' - Dickie's bladderfern *'' Cystopteris douglasii'' - Douglas' bladderfern *''Cystopteris fragilis'' - ...
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Cystopteris Protrusa
''Cystopteris protrusa'' is a common fern of North America, commonly known as the lowland bladderfern, lowland brittle fern or lowland fragile fern. The plant is native to eastern Canada, and the Midwestern and Eastern United States. Throughout much of its range it is the most common ''Cystopteris'' fern species. Description Unlike most species of the genus ''Cystopteris'', this fern is exclusively terrestrial, often forming large, dense colonies. It is also largely a spring ephemeral. Some fronds may remain by late summer, but most have disappeared. The specific name, protrusa, refers to the fact that the rhizome protrudes a short distance beyond the current year's fronds to form the following year's leaf buds. This protrusion is a positive diagnostic in the field. Sori are round and covered in a bladder-like indusium. Taxonomy ''Cystopteris protrusa'' was originally considered only a variety of ''Cystopteris fragilis''. However, it is now known that this is a diploid s ...
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Cystopteris Bulbifera
''Cystopteris bulbifera'', with the common name bulblet fern, bulblet bladderfern, or bulblet fragile fern is a fern in the family Cystopteridaceae. Distribution The fern is native to eastern Canada, the Midwestern and Eastern United States, and two disjunct populations in the Southwestern United States. It is found only on calcareous substrates such as limestone. It commonly festoons limestone cave openings. While most commonly found on vertical rock faces, it also grows in rocky scree. Description ''Cystopteris bulbifera'' is a low-growing rock fern with creeping stems and narrow elongate deltate fronds which grow to 75 cm (30 in). ''C. bulbifera'' is unusual among ferns in producing bulblets along the bottom of the fronds. This is one of the easiest ''Cystopteris'' species to identify. Hybrids This species is known to hybridize with ''Cystopteris fragilis'', ''Cystopteris protrusa'', ''Cystopteris reevesiana'', and ''Cystopteris tenuis''. Hybrids with ''C. fragil ...
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Cystopteris Tenuis
''Cystopteris tenuis'' is sometimes known as Mackay's bladder fern or Mackay's fragile fern. It was long considered to be a part of the superspecies for fragile ferns, as ''Cystopteris fragilis'' ( L.) Bernh. var. ''mackayi'' Lawson. left, ''C. tenuis'' in native habitat This species is an allotetraploid of hybrid origin (see Cystopteris hybrid complex). The parent diploid species are ''Cystopteris protrusa'' and the hypothesized ''Cystopteris hemifragilis'', believed to be an extinct species. ''C. tenuis'' is known to hybridize with '' C. bulbifera'' to produce the hybrid ''C. Xillinoensis'', with '' C. tennesseensis'' top produce the hybrid ''C. Xwagneri'', and with '' C. fragilis'' and '' C. protrusa'' to produce unnamed hybrids (as per Flora of North America). Mackay's fragile fern grows on rock or in scree, generally in sheltered spots, in the northeastern United States. It may be distinguished from the somewhat similar ''Cystopteris tennesseensis ''Cystopteris'' ...
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Cystopteris Dickieana
''Cystopteris dickieana'', commonly known as Dickie's bladder-fern, is a fern with a wide distribution in the Northern Hemisphere. There is debate amongst botanists as to whether it is a species in its own right or a variant of '' C. fragilis''. Distribution ''C. dickieana'' is native to Canada and the United States, a variety of European countries including Russia and also north Africa and the Andes. It is typically found in montane habitats below the tree-line, although it is also grows at lower altitudes in locations with cool summers. Discovery and Victorian collectors The first recorded discovery of the plant was made by William Knight, Professor of Natural Philosophy at Marischal College, Aberdeen in Scotland. Knight came across a small population growing on base-rich rocks in a sea cave (known locally as a "yawn") on the coast of Kincardineshire. The first publication to record it was the 1838 ''Flora Aberdonenis'' which included a note of its occurrence written by a pup ...
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Cystopteris Montana
''Cystopteris montana,'' previously classified as '' Athyrium montanum'', is a species of fern known by the common name mountain bladderfern. It occurs throughout the high latitudes of the Northern Hemisphere, in Eurasia, Greenland, and Alaska, and throughout Canada. It is also present in the higher elevations in Colorado farther south.''Cystopteris montana''.
Flora of North America.
This fern produces a creeping, cordlike, scaly stem. The leaves are up to 45 centimeters long. The blades are borne on a petiole with a dark base and a light-colored end. The petiole is longer than the blade. The blade is



Cystopteris Alpina
''Cystopteris alpina'' is a fern in the family Cystopteridaceae. It is closely related to '' C. fragilis'' and has been treated as conspecific with that species by many authors.Jonsell B (ed). (2000). "Flora Nordica: Volume 1 - Lycopodiacea to Polygonaceae".Bergius Foundation, Stockholm However, according to the Flora of North America, it is an allopolyploid species of hybrid origin, with ''Cystopteris montana'' as one probable parent. It is known to hybridise with '' C. fragilis'' in Scandinavia and intermediate plants possibly of hybrid origin are known from North Wales.Tennant DJ. (2010). “The British records of ''Cystopteris alpina'' (Lamarck) Desvaux; Woodsiaceae”. ''Watsonia'' 28: 57-63 Distribution ''C. alpina'' is a montane species occurring through most of Europe with populations extending into Russia and West-Asian mountain regions. It is widely distributed through Northern Scandinavia, the Alps and the Pyrenees. In Britain it is thought to be extinct having only be ...
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