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Cyathea Holdridgeana
''Cyathea holdridgeana'', synonym ''Cyathea albomarginata'', is a species of tree fern native to Panamá and Costa Rica. It grows in wet forests at elevations of 2400–2800 m, considerably higher than most other tree ferns of Central America. ''Cyathea holdridgeana'' is a small tree with a trunk up to 50 cm tall. Petiole purplish-brown with white margins, scaly but not spiny; scales (= modified appendages attached to the margins of the petioles) bicolored, dark purple in the center but white along the margins; dead scales appearing totally white. Leaves bipinnate The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular o ...; pinnulae 10–12 cm long References holdridgeana Flora of Costa Rica Flora of Panama {{Cyatheales-stub ...
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Synonym (taxonomy)
The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called ''Pinus abies''. This name is no longer in use, so it is now a synonym of the current scientific name, ''Picea abies''. * In zoology, moving a species from one genus to another results in a different binomen, but the name is considered an alternative combination rather than a synonym. The concept of synonymy in zoology is reserved for two names at the same rank that refers to a taxon at that rank - for example, the name ''Papilio prorsa'' Linnaeus, 1758 is a junior synonym of ''Papilio levana'' Linnaeus, 1758, being names for different seasonal forms of the species now referred to as ''Araschnia le ...
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Tree Fern
The tree ferns are arborescent (tree-like) ferns that grow with a trunk elevating the fronds above ground level, making them trees. Many extant tree ferns are members of the order Cyatheales, to which belong the families Cyatheaceae (scaly tree ferns), Dicksoniaceae, Metaxyaceae, and Cibotiaceae. It is estimated that Cyatheales originated in the early Jurassic, and is the third group of ferns known to have given rise to tree-like forms. The others are the extinct '' Tempskya'' of uncertain position, and Osmundales where the extinct Guaireaceae and some members of Osmundaceae also grew into trees. In addition there were the Psaroniaceae and Tietea in the Marattiales, which is the sister group to most living ferns including Cyatheales. Other ferns which are also tree ferns, are ''Leptopteris'' and ''Todea'' in the family Osmundaceae, which can achieve short trunks under a metre tall. Fern species with short trunks in the genera ''Blechnum'', '' Cystodium'' and ''Sadleria'' from ...
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Panamá
Panama ( , ; es, link=no, Panamá ), officially the Republic of Panama ( es, República de Panamá), is a transcontinental country spanning the southern part of North America and the northern part of South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and the Pacific Ocean to the south. Its capital and largest city is Panama City, whose metropolitan area is home to nearly half the country's million people. Panama was inhabited by indigenous tribes before Spanish colonists arrived in the 16th century. It broke away from Spain in 1821 and joined the Republic of Gran Colombia, a union of Nueva Granada, Ecuador, and Venezuela. After Gran Colombia dissolved in 1831, Panama and Nueva Granada eventually became the Republic of Colombia. With the backing of the United States, Panama seceded from Colombia in 1903, allowing the construction of the Panama Canal to be completed by the United States Army Corps of Engin ...
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Costa Rica
Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, the Pacific Ocean to the southwest, and Maritime boundary, maritime border with Ecuador to the south of Cocos Island. It has a population of around five million in a land area of . An estimated 333,980 people live in the capital and largest city, San José, Costa Rica, San José, with around two million people in the surrounding metropolitan area. The sovereign state is a Unitary state, unitary Presidential system, presidential Constitution of Costa Rica, constitutional republic. It has a long-standing and stable democracy and a highly educated workforce. The country spends roughly 6.9% of its budget (2016) on education, compared to a global average of 4.4%. Its economy, once heavily dependent on agricultu ...
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Petiole (botany)
In botany, the petiole () is the stalk that attaches the leaf blade to the stem, and is able to twist the leaf to face the sun. This gives a characteristic foliage arrangement to the plant. Outgrowths appearing on each side of the petiole in some species are called stipules. Leaves with a petiole are said to be petiolate, while leaves lacking a petiole are called sessile or apetiolate. Description The petiole is a stalk that attaches a leaf to the plant stem. In petiolate leaves, the leaf stalk may be long, as in the leaves of celery and rhubarb, or short. When completely absent, the blade attaches directly to the stem and is said to be sessile. Subpetiolate leaves have an extremely short petiole, and may appear sessile. The broomrape family Orobanchaceae is an example of a family in which the leaves are always sessile. In some other plant groups, such as the speedwell genus '' Veronica'', petiolate and sessile leaves may occur in different species. In the grasses (Poaceae), ...
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Bipinnate
The following is a list of terms which are used to describe leaf morphology in the description and taxonomy of plants. Leaves may be simple (a single leaf blade or lamina) or compound (with several leaflets). The edge of the leaf may be regular or irregular, may be smooth or bearing hair, bristles or spines. For more terms describing other aspects of leaves besides their overall morphology see the leaf article. The terms listed here all are supported by technical and professional usage, but they cannot be represented as mandatory or undebatable; readers must use their judgement. Authors often use terms arbitrarily, or coin them to taste, possibly in ignorance of established terms, and it is not always clear whether because of ignorance, or personal preference, or because usages change with time or context, or because of variation between specimens, even specimens from the same plant. For example, whether to call leaves on the same tree "acuminate", "lanceolate", or "linear" could ...
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Pinnule
A leaflet (occasionally called foliole) in botany is a leaf-like part of a compound leaf. Though it resembles an entire leaf, a leaflet is not borne on a main plant stem or branch, as a leaf is, but rather on a petiole or a branch of the leaf. Compound leaves are common in many plant families and they differ widely in morphology. The two main classes of compound leaf morphology are palmate and pinnate. For example, a ''hemp'' plant has palmate compound leaves, whereas some species of ''Acacia'' have pinnate leaves. The ultimate free division (or leaflet) of a compound leaf, or a pinnate subdivision of a multipinnate leaf is called a pinnule or pinnula. Image:Ветвь акации.jpg, Pinnate leaf of a legume with 10 leaflets Image:Mimosa Pudica.gif, ''Mimosa pudica'' folding leaflets inward. See also * Compound leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosyn ...
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Cyathea
''Cyathea'' is a genus of tree ferns, the type genus of the fern order Cyatheales. The genus name ''Cyathea'' is derived from the Greek ''kyatheion'', meaning "little cup", and refers to the cup-shaped sori on the underside of the fronds. Description The species of ''Cyathea'' are mostly terrestrial ferns, usually with a single tall stem. Rarely, the trunk may be branched or creeping. Many species also develop a fibrous mass of roots at the base of the trunk. The genus has a pantropical distribution, with over 470 species. They grow in habitats ranging from tropical rain forests to temperate woodlands. Classification Conant ''et al.'' in 1996, concluded on molecular cpDNA and morphological evidence that a system of three clades – ''Alsophila'', ''Cyathea'' and ''Sphaeropteris'' was the most accurate reflection of evolutionary lineages within the Cyatheaceae, ''Alsophila'' being the most basal and ''Cyathea'' and ''Sphaeropteris'' derived sister groups. In the Pteridophyte ...
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Flora Of Costa Rica
Costa Rica (, ; ; literally "Rich Coast"), officially the Republic of Costa Rica ( es, República de Costa Rica), is a country in the Central American region of North America, bordered by Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, the Pacific Ocean to the southwest, and maritime border with Ecuador to the south of Cocos Island. It has a population of around five million in a land area of . An estimated 333,980 people live in the capital and largest city, San José, with around two million people in the surrounding metropolitan area. The sovereign state is a unitary presidential constitutional republic. It has a long-standing and stable democracy and a highly educated workforce. The country spends roughly 6.9% of its budget (2016) on education, compared to a global average of 4.4%. Its economy, once heavily dependent on agriculture, has diversified to include sectors such as finance, corporate services for foreign companies, pharma ...
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