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Ctenosauriscids
Ctenosauriscidae is an extinct family of pseudosuchian archosaurs within the clade Poposauroidea. Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest Olenekian to Anisian stages). All species had large "sails" on their backs. Ctenosauriscids are among some of the earliest archosaurs and represent the first global radiation of the group. Genera Phylogeny Ctenosauriscidae was named by Oskar Kuhn in 1964 to include the genus ''Ctenosauriscus''. It is a stem-based taxon defined by Richard J. Butler, Stephen L. Brusatte, Mike Reich, Sterling J. Nesbitt, Rainer R. Schoch and Jahn J. Hornung in 2011 as "the most inclusive clade containing ''Ctenosauriscus koeneni'' but not ''Poposaurus gracilis'', ''Effigia okeeffeae'', ''Postosuchus kirkpatricki'', ''Crocodylus niloticus'', '' Ornithosuchus longidens'', or '' Aetosaurus ferratus''". The cladogram A cladogram (from Greek ''clados'' "branch" and ''gramm ...
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Arizonasaurus
''Arizonasaurus'' was a ctenosauriscid archosaur from the Middle Triassic (243 million years ago). ''Arizonasaurus'' is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947. Discovery and naming The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947 on the basis of a few teeth and a maxilla, labelled as specimen UCMP 36232. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. Description ''Arizonasaurus'' had a sail made of tall neural spines. This sail was similar to those of other basal archosaurs, such as other ctenosauriscids like ''Ctenosauriscus'', ''Lotosaurus'', ''Bromsgroveia'', and ''Hypselorhachis''. ''Arizonasaurus'' is described from two braincase specimens. Some ancestral f ...
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Bromsgroveia
''Bromsgroveia'' is an extinct genus of predatory ctenosauriscid archosaur from the Middle Triassic Bromsgrove Sandstone of England. Ctenosauriscids were a group of rauisuchians that was related to the ancestors of modern crocodiles and alligators. Classification ''Bromsgroveia'' was closely related to ''Ctenosauriscus'', and together with a few other genera they make up Ctenosauriscidae Ctenosauriscidae is an extinct family (biology), family of pseudosuchian archosaurs within the clade Poposauroidea. Ctenosauriscids existed in Africa, Asia, Europe and North America during the Early Triassic to the Middle Triassic period (latest .... The ctenosauriscids were closely related to the Poposauridae, poposaurids, as shown by a few shared derived characteristics. The pelvic girdle in ''Bromsgroveia'' unites this taxon with ''Ctenosauriscus'', ''Lotosaurus'', ''Arizonasaurus'', and ''Hypselorhachus''. Below is a phylogenetic cladogram simplified from Butler ''et al.'' in 2011 showing ...
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Arizonasaurus BW
''Arizonasaurus'' was a Ctenosauriscidae, ctenosauriscid archosaur from the Middle Triassic (243 million years ago). ''Arizonasaurus'' is found in the Middle Triassic Moenkopi Formation of northern Arizona. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. The taxon has a large sailback formed by elongated neural spines of the vertebrae. The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947. Discovery and naming The type species, ''Arizonasaurus babbitti'', was named by Samuel Paul Welles in 1947 on the basis of a few teeth and a maxilla, labelled as specimen UCMP 36232. A fairly complete skeleton was found in 2002 by Sterling Nesbitt. Description ''Arizonasaurus'' had a sail made of tall neural spines. This sail was similar to those of other basal archosaurs, such as other ctenosauriscidae, ctenosauriscids like ''Ctenosauriscus'', ''Lotosaurus'', ''Bromsgroveia'', and ''Hypselorhachis''. ''Arizonasaurus'' is described from two br ...
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Xilousuchus
''Xilousuchus'' is an extinct genus of poposauroid from lower Triassic (Olenekian stage) deposits of Fugu County of northeastern Shanxi Province, China. It is known from the holotype, IVPP V 6026, a single well-preserved partial skeleton including the skull. It was found from the Heshanggou Formation of the Ordos Basin, Hazhen commune. It was first named by Xiao-Chun Wu in 1981 and the type species is ''Xilousuchus sapingensis''. Wu (1981) referred ''Xilousuchus'' to the Proterosuchia. Gower and Sennikov (1996) found it to be an erythrosuchian based strictly on the braincase. A more detailed re-description of the genus was provided by Nesbitt ''et al.'' (2010) and found poposauroid affinities. In his massive revision of archosaurs which included a large cladistic analysis, Sterling J. Nesbitt (2011) found ''Xilousuchus'' to be a poposauroid which is most closely related to ''Arizonasaurus''. ''Xilousuchus'' is the oldest archosaur to date, although ''Ctenosauriscus'' ...
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Ctenosauriscus Koeneni
''Ctenosauriscus'' is an extinct genus of sail-backed poposauroid archosaur from Early Triassic deposits of Lower Saxony in northern Germany. It gives its name to the family Ctenosauriscidae, which includes other sail-backed poposauroids such as ''Arizonasaurus''. Fossils have been found in latest Olenekian deposits around 247.5-247.2 million years old, making it one of the first known archosaurs. History of discovery ''Ctenosauriscus'' is known from the holotype, GZG.V.4191, a single partial preserved postcranial skeleton including partial vertebral column, ribs and girdle. The holotype is composed of four slabs, which were labeled A1, A2, B1, and B2 in a 2011 study. Slabs A1 and B1 form the part and slabs A2 and B2 form the counterpart. The holotype was discovered in early 1871 in the Bremke dell locality, near the county of Göttingen. The holotype was found in a deposit called the Solling-Bausandstein, which is an outcropping of the upper Middle Buntsandstein in the region of ...
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Bystrowisuchus
''Bystrowisuchus'' is an extinct genus of ctenosauriscid pseudosuchian archosaur from the Early Triassic of European Russia. Fossils have been found in the Olenekian-age Lipovskaya Formation in Ilovlinsky District. The type species is ''Bystrowisuchus flerovi''. Description ''Bystrowisuchus flerovi'' is based on a holotype specimen including six cervical or neck vertebrae and a partial right ilium or hip bone. Its total body length is estimated at . Discovery The holotype specimen of ''Bystrowisuchus'' was found near the eastern banks of the Don River in Ilovlinsky District, Volgograd Oblast. It came from a fossil site known as the Donskaya Luka locality, which preserves a wide diversity of Early Triassic tetrapod fossils. Along with ''Bystrowisuchus'', the Donskaya Luka fossil assemblage includes temnospondyl amphibians, a chroniosuchian, a procolophonid, a sauropterygian, a protorosaurian, possible trilophosaurid archosauromorphs, two rauisuchid archosaurs, and a dicynodont ...
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Ctenosauriscus
''Ctenosauriscus'' is an extinct genus of sail-backed poposauroid archosaur from Early Triassic deposits of Lower Saxony in northern Germany. It gives its name to the family Ctenosauriscidae, which includes other sail-backed poposauroids such as ''Arizonasaurus''. Fossils have been found in latest Olenekian deposits around 247.5-247.2 million years old, making it one of the first known archosaurs. History of discovery ''Ctenosauriscus'' is known from the holotype, GZG.V.4191, a single partial preserved postcranial skeleton including partial vertebral column, ribs and girdle. The holotype is composed of four slabs, which were labeled A1, A2, B1, and B2 in a 2011 study. Slabs A1 and B1 form the part and slabs A2 and B2 form the counterpart. The holotype was discovered in early 1871 in the Bremke dell locality, near the county of Göttingen. The holotype was found in a deposit called the Solling-Bausandstein, which is an outcropping of the upper Middle Buntsandstein in the region of ...
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Hypselorhachis
''Hypselorhachis'' is a genus of extinct reptile, possibly a ctenosauriscid archosaur related to ''Ctenosauriscus''. It lived during the Triassic Period. It is currently known only from a single vertebra found from the Middle Triassic Manda Beds in Tanzania. The vertebra is preserved in reasonably good condition, as although the tall neural spine is chipped in several places it is not broken despite being quite slender, only around 20 mm thick transversely. The type species is ''H. mirabilis'', mentioned but never fully described by English paleontologist Alan J. Charig. ''Hypselorhachis'' was assigned to the Ctenosauriscidae, a group of sail-backed archosaurs, in 1988.Carroll, R.L. (1988). ''Vertebrate Paleontology and Evolution''. W.H. Freeman and Company It was formally described by Richard J. Butler and co-workers in 2009. The name means 'wonderful high spine', from the Latin 'mirabilis' 'wonderful' and the Greek 'ὑπσελος', 'high' and 'ῥαχις' 'spine' or ...
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Ctenosauriscus BW
''Ctenosauriscus'' is an extinct genus of sail-backed poposauroid archosaur from Early Triassic deposits of Lower Saxony in northern Germany. It gives its name to the family Ctenosauriscidae, which includes other sail-backed poposauroids such as ''Arizonasaurus''. Fossils have been found in latest Olenekian deposits around 247.5-247.2 million years old, making it one of the first known archosaurs. History of discovery ''Ctenosauriscus'' is known from the holotype, GZG.V.4191, a single partial preserved postcranial skeleton including partial vertebral column, ribs and girdle. The holotype is composed of four slabs, which were labeled A1, A2, B1, and B2 in a 2011 study. Slabs A1 and B1 form the part and slabs A2 and B2 form the counterpart. The holotype was discovered in early 1871 in the Bremke dell locality, near the county of Göttingen. The holotype was found in a deposit called the Solling-Bausandstein, which is an outcropping of the upper Middle Buntsandstein in the region of ...
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Solling Formation
The Solling Formation is a geologic formation in Germany.Solling Formation
at .org
Formerly considered earliest (), but later dating places the formation from the to

Bromsgrove Sandstone Formation
The Sherwood Sandstone Group is a Triassic lithostratigraphic group (a sequence of rock strata) which is widespread in Britain, especially in the English Midlands. The name is derived from Sherwood Forest in Nottinghamshire which is underlain by rocks of this age. It has economic importance as the reservoir of the Morecambe Bay gas field, the second largest gas field in the UK. Geographical extent These rocks are found in northwest England as far north as Carlisle (and extending just into Scotland around Annan and Gretna) and in the Vale of Eden and then extending down the Cumbrian coast into Lancashire and Cheshire. They are mostly obscured by superficial deposits but the highest coastal cliffs in northwest England at St Bees Head are formed in the St Bees Sandstone, the lowermost formation within the group. In the northeast they extend from Hartlepool south through the Vale of Mowbray and the Vale of York, then south through Nottinghamshire into the English Midlands, though ar ...
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Lipovskaya Formation
The Yarenskian Gorizont (geology), Gorizont ("Yarenskian Horizon") is a major biostratigraphic unit in Russia corresponding to Upper Olenekian-age terrestrial sediments. It lies above the Ustmylian Gorizont of the Vetlugian Supergorizont, and is located below a short unconformity overlain by the Donguz Gorizont. Some paleontologists elevate the Yarenskian to a supergorizont and subdivide it into two smaller units: the older Fedorovskian Gorizont and the younger Gamskian Gorizont. Others prefer to consider the Fedorovskian and Gamskian to be subgorizonts or members of the Yarenskian. Vertebrate index fossils of the Fedorovskian include the lungfish ''Gnathorhiza,'' the Trematosauridae, trematosaurid ''Inflectosaurus,'' and the procolophonid ''Burtensia''. In the Gamskian, these index fossils are replaced by ''Ceratodus'', ''Trematosaurus'', and ''Kapes (genus), Kapes'', respectively. The Yarenskian as a whole is characterized by the capitosaur amphibian ''Parotosuchus'', and has also ...
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