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Cruziana
''Cruziana'' is a trace fossil consisting of elongate, bilobed, approximately bilaterally symmetrical burrows, usually preserved along bedding planes, with a sculpture of repeated striations that are mostly oblique to the long dimension. It is found in marine and freshwater sediments. It first appears in upper Fortunian rocks of northern Iran and northern Norway. ''Cruziana'' has been extensively studied because it has uses in biostratigraphy (specific scratch patterns are unique to specific time intervals), and because the traces can reveal many aspects of their makers' behavior. ''Cruziana'' is typically associated with trilobites but can also made by other arthropods. ''Cruziana'' appears in non-marine formations such as the Beacon Supergroup that would have been unsuitable environments for trilobites, and in Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beg ...
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Cruziana Brallier Harrell Devonian
''Cruziana'' is a trace fossil consisting of elongate, bilobed, approximately bilaterally symmetrical burrows, usually preserved along bedding planes, with a sculpture of repeated striations that are mostly oblique to the long dimension. It is found in marine and freshwater sediments. It first appears in upper Fortunian rocks of northern Iran and northern Norway. ''Cruziana'' has been extensively studied because it has uses in biostratigraphy (specific scratch patterns are unique to specific time intervals), and because the traces can reveal many aspects of their makers' behavior. ''Cruziana'' is typically associated with trilobites but can also made by other arthropods. ''Cruziana'' appears in non-marine formations such as the Beacon Supergroup that would have been unsuitable environments for trilobites, and in Triassic sediments that were deposited after trilobites became extinct at the Permian-Triassic extinction event, end of the Permian Period. ''Cruziana'' traces can reach ...
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Trace Fossil
A trace fossil, also known as an ichnofossil (; from el, ἴχνος ''ikhnos'' "trace, track"), is a fossil record of biological activity but not the preserved remains of the plant or animal itself. Trace fossils contrast with body fossils, which are the fossilized remains of parts of organisms' bodies, usually altered by later chemical activity or mineralization. The study of such trace fossils is ichnology and is the work of ichnologists. Trace fossils may consist of impressions made on or in the substrate by an organism. For example, burrows, borings (bioerosion), urolites (erosion caused by evacuation of liquid wastes), footprints and feeding marks and root cavities may all be trace fossils. The term in its broadest sense also includes the remains of other organic material produced by an organism; for example coprolites (fossilized droppings) or chemical markers (sedimentological structures produced by biological means; for example, the formation of stromatolites). H ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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Trilobites
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stud ...
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Rusophycus
''Rusophycus'' is a trace fossil ichnogenus allied to ''Cruziana''. ''Rusophycus'' is the resting trace, recording the outline of the tracemaker; ''Cruziana'' is made when the organism moved. The sculpture of ''Rusophycus'' may reveal the approximate number of legs that the tracemaker had, although striations (scratchmarks) from a single leg may overlap or be repeated. Both ''Rusophycus'' and ''Cruziana'' are typically associated with trilobites but can also be made by other arthropods. ''Rusophycus'' is known from sediments dating to the earliest Cambrian Period (somewhat before the first appearance trilobite fossils), and from the Triassic Period (after the Permian The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last ... extinction of trilobites), indicating that other producers cou ...
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Beacon Supergroup
The Beacon Supergroup is a geological formation exposed in Antarctica and deposited from the Devonian to the Triassic (). The unit was originally described as either a formation or sandstone, and upgraded to group and supergroup as time passed. It contains a sandy member known as the Beacon Heights Orthoquartzite. Overview The base of the Beacon Supergroup is marked by an unconformity and is composed of the Devonian Taylor Group, a quartzose sandstone sequence; and the Late Carboniferous to Early Jurassic Victoria Group, consisting of glacial beds, sandstone, shale, conglomerate, and coal. The Beacon Sandstone was named by Hartley T. Ferrar during Scott's Discovery Expedition (1901–1904), using the Beacon Heights survey points as reference. ''Glossopteris'' fossils dated the sandstone to the Permian and linked the lithology to similar sequences on neighboring continents. Generally flat lying, the supergroup is up to 3.2 km thick and is fairly continuous from south Victor ...
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Diplichnites
''Diplichnites'' are arthropod Fossil trackway, trackways with two parallel rows of blunt to elongate, closely spaced tracks oriented approximately perpendicularly to the mid-line of the trackway. The term is more often used for the ichnofossils of this description; however, similar trackways from recent arthropods are sometimes given this name as well. The ''Diplichnites'' makers This type of trackway was originally based on large fossils from Pennsylvanian (geology), Pennsylvanian strata of Nova Scotia, when Sir J. W. Dawson named it in 1873. Dawson proposed that ''Diplichnites'' were produced by a fish “walking” in shallow water on pectoral or ventral fin spines. Previous to this he had suggested that a large crustacean, annelid worm or myriapod (such as a millipede) could have made them. Subsequent evidence has supported this earlier interpretation. The fossils on which this ichnotaxon was based are now thought to be from giant myriapods, such as ''Arthropleura''. The ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Triassic
The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest period of the Mesozoic Era. Both the start and end of the period are marked by major extinction events. The Triassic Period is subdivided into three epochs: Early Triassic, Middle Triassic and Late Triassic. The Triassic began in the wake of the Permian–Triassic extinction event, which left the Earth's biosphere impoverished; it was well into the middle of the Triassic before life recovered its former diversity. Three categories of organisms can be distinguished in the Triassic record: survivors from the extinction event, new groups that flourished briefly, and other new groups that went on to dominate the Mesozoic Era. Reptiles, especially archosaurs, were the chief terrestrial vertebrates during this time. A specialized subgroup of archo ...
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Fortunian
The Fortunian age marks the beginning of the Phanerozoic Eon, the Paleozoic Era, and the Cambrian Period. It is the first of the two stages of the Terreneuvian series. Its base is defined as the first appearance of the trace fossil ''Treptichnus pedum'' million years ago. The top of the Fortunian which is the base of the Stage 2 of the Cambrian has not been formally defined yet, but will correspond to the appearance of an Archeocyatha species or "Small shelly fossils" approximately million years ago. The name Fortunian is derived from a part of the Burin Peninsula, the town of Fortune near the GSSP and Fortune Bay. GSSP The type locality (GSSP) of the Fortunian stage is in Fortune Head, at the northern edge of the Burin Peninsula, Newfoundland, Canada (). This GSSP coincides with the base of the Terreneuvian series, the Precambrian-Cambrian boundary and the beginning of the Phanerozoic. The outcrops show a carbonate-siliciclastic succession which is mapped as the Chapel Island F ...
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Biostratigraphy
Biostratigraphy is the branch of stratigraphy which focuses on correlating and assigning relative ages of rock Stratum, strata by using the fossil assemblages contained within them.Hine, Robert. “Biostratigraphy.” ''Oxford Reference: Dictionary of Biology'', 8th ed., Oxford University Press, 2019. The primary objective of biostratigraphy is ''correlation'', demonstrating that a particular Horizon (geology), horizon in one geological section represents the same period of time as another horizon at a different section. Fossils within these strata are useful because sediments of the same age can look completely different, due to local variations in the Sedimentary depositional environment, sedimentary environment. For example, one section might have been made up of clays and marls, while another has more chalky limestones. However, if the fossil species recorded are similar, the two sediments are likely to have been laid down around the same time. Ideally these fossils are used to ...
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Arthropods
Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a Segmentation (biology), segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and Arthropod cuticle, cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate. The arthropod body plan consists of segments, each with a pair of appendages. Arthropods are bilaterally symmetrical and their body possesses an exoskeleton, external skeleton. In order to keep growing, they must go through stages of moulting, a process by which they shed their exoskeleton to reveal a new one. Some species have wings. They are an extremely diverse group, with up to 10 million species. The haemocoel, an arthropod's internal cavity, through which its haemolymph – analogue of blood – circulates, accommodates its interior Organ (anatomy), organs; it has an open circulatory system. Like their exteriors, the internal or ...
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