Cruralispennia
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Cruralispennia
''Cruralispennia'' is an extinct genus of enantiornithean bird. The only known specimen of ''Cruralispennia'' was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of ''Cruralispennia'' is ''Cruralispennia multidonta''. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of ''Cruralispennia'' is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers. ''Cruralispennia'' lived alongside several other species of enantiornitheans, such as ''Protopteryx,'' in the Sichakou Basin of the Huajiying Formation, approximately 130.7 million years ago. Despite being one of the earliest known ornithothoraces, this genus possessed several features of derived enantiornitheans as well as some unusual traits convergent with ornithuromorphs, a group containing modern birds. ''Cruralispennia'' is also notable for having crural feathers or ...
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Cruralispennia Reconstruction
''Cruralispennia'' is an extinct genus of enantiornithean bird. The only known specimen of ''Cruralispennia'' was discovered in the Early Cretaceous Huajiying Formation of China and formally described in 2017. The type species of ''Cruralispennia'' is ''Cruralispennia multidonta''. The generic name is Latin for "shin feather", while the specific name means "many-toothed". The holotype of ''Cruralispennia'' is IVPP 21711, a semi-articulated partial skeleton surrounded by the remains of carbonized feathers. ''Cruralispennia'' lived alongside several other species of enantiornitheans, such as ''Protopteryx,'' in the Sichakou Basin of the Huajiying Formation, approximately 130.7 million years ago. Despite being one of the earliest known ornithothoraces, this genus possessed several features of derived enantiornitheans as well as some unusual traits convergent with ornithuromorphs, a group containing modern birds. ''Cruralispennia'' is also notable for having crural feathers origina ...
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Enantiornithes
The Enantiornithes, also known as enantiornithines or enantiornitheans in literature, are a group of extinct avialans ("birds" in the broad sense), the most abundant and diverse group known from the Mesozoic era. Almost all retained teeth and clawed fingers on each wing, but otherwise looked much like modern birds externally. Over eighty species of Enantiornithes have been named, but some names represent only single bones, so it is likely that not all are valid. The Enantiornithes became extinct at the Cretaceous–Paleogene boundary, along with Hesperornithes and all other non-avian dinosaurs. Discovery and naming The first Enantiornithes to be discovered were incorrectly referred to modern bird groups. For example, the first known species of Enantiornithes, ''Gobipteryx minuta'', was originally considered a paleognath related to ostriches and tinamou. The Enantiornithes were first recognized as a distinct lineage, or "subclass" of birds, by Cyril A. Walker in 1981. Walker mad ...
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Huajiying Formation
The Huajiying Formation is a geological formation in Hebei, People's Republic of China. Known for its fossils including feathered dinosaurs, the age of the formation is uncertain. It may represent an early portion of the Jehol Biota, dating to somewhere in the early Cretaceous or late Jurassic periods. It may correlate with the early Cretaceous Dadianzi Formation and parts of the Yixian Formation, with an age range between 140 and 122 Ma ago.Wang, X., O’Connor, J.K., Zhao, B., Chiappe, L.M., Gao, C. and Cheng, X. (2010). "New species of Enantiornithes (Aves: Ornithothoraces) from the Qiaotou Formation in Northern Hebei, China." ''Acta Geologica Sinica'', 84(2): 247-256. It contains the Qiaotou Member, sometimes treated as a distinct formation. Age and stratigraphy The Huajiying Formation contains the following members, in order from youngest to oldest: * 5th Volcanic Member * Guohedao Sedimentary Member * 4th Volcanic Member * Nianzigou Sedimentary Member * 3rd Volcanic Member ...
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Ornithothoraces
Ornithothoraces is a group of avialans that includes all enantiornithes ("opposite birds") and the euornithes ("true birds"), which includes modern birds and their closest ancestors. The name Ornithothoraces means "bird thoraxes". This refers to the modern, highly advanced anatomy of the thorax that gave the ornithothoracines superior flight capability compared with more primitive avialans. This anatomy includes a large, keeled breastbone, elongated coracoids and a modified glenoid joint in the shoulder, and a semi-rigid rib cage. In spite of this at least the sternum seems to have developed convergently rather than being a true homology. The earliest known members of the group are the enantiornitheans ''Protopteryx fengningensis'', '' Eopengornis martini'', and '' Cruralispennia multidonta'', as well as the euornithine '' Archaeornithura meemannae'', all from the Sichakou Member of the Huajiying Formation in China, which has been dated to 130.7 million years old. At least one ot ...
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Confuciusornithidae
Confuciusornithidae is an extinct family of pygostylian avialans known from the Early Cretaceous, found in northern China. They are commonly placed as a sister group to Ornithothoraces, a group that contains all extant birds along with their closest extinct relatives. Confuciusornithidae contains four genera, possessing both shafted and non-shafted (downy) feathers. They are also noted for their distinctive pair of ribbon-like tail feathers of disputed function. The wing anatomy of confuciusornithids suggests an unusual flight behavior, due to anatomy that implies conflicting abilities. They possessed feathers similar to those of fast-flapping birds, which rely on quick flapping of their wings to stay aloft. At the same time, their wing anatomy also suggests a lack of flapping ability. Confuciusornithids are also noted for their beak and lack of teeth, similar to modern birds. Both predators and prey, confuciusornithid fossils have been observed with fish remains in their digestive ...
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Premaxilla
The premaxilla (or praemaxilla) is one of a pair of small cranial bones at the very tip of the upper jaw of many animals, usually, but not always, bearing teeth. In humans, they are fused with the maxilla. The "premaxilla" of therian mammal has been usually termed as the incisive bone. Other terms used for this structure include premaxillary bone or ''os premaxillare'', intermaxillary bone or ''os intermaxillare'', and Goethe's bone. Human anatomy In human anatomy, the premaxilla is referred to as the incisive bone (') and is the part of the maxilla which bears the incisor teeth, and encompasses the anterior nasal spine and alar region. In the nasal cavity, the premaxillary element projects higher than the maxillary element behind. The palatal portion of the premaxilla is a bony plate with a generally transverse orientation. The incisive foramen is bound anteriorly and laterally by the premaxilla and posteriorly by the palatine process of the maxilla. It is formed from the ...
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Frontal Bone
The frontal bone is a bone in the human skull. The bone consists of two portions.''Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, part of the bony orbital cavity holding the eye, and part of the bony part of the nose respectively. The name comes from the Latin word ''frons'' (meaning " forehead"). Structure of the frontal bone The frontal bone is made up of two main parts. These are the squamous part, and the orbital part. The squamous part marks the vertical, flat, and also the biggest part, and the main region of the forehead. The orbital part is the horizontal and second biggest region of the frontal bone. It enters into the formation of the roofs of the orbital and nasal cavities. Sometimes a third part is included as the nasal part of the frontal bone, and sometimes this is included with the squamous part. The nasal part is between the brow ridges, and ends in ...
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Dentary
In anatomy, the mandible, lower jaw or jawbone is the largest, strongest and lowest bone in the human facial skeleton. It forms the lower jaw and holds the lower tooth, teeth in place. The mandible sits beneath the maxilla. It is the only movable bone of the skull (discounting the ossicles of the middle ear). It is connected to the temporal bones by the temporomandibular joints. The bone is formed prenatal development, in the fetus from a fusion of the left and right mandibular prominences, and the point where these sides join, the mandibular symphysis, is still visible as a faint ridge in the midline. Like other symphyses in the body, this is a midline articulation where the bones are joined by fibrocartilage, but this articulation fuses together in early childhood.Illustrated Anatomy of the Head and Neck, Fehrenbach and Herring, Elsevier, 2012, p. 59 The word "mandible" derives from the Latin word ''mandibula'', "jawbone" (literally "one used for chewing"), from ''wikt:mandere ...
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Pygostyle
Pygostyle describes a skeletal condition in which the final few caudal vertebrae are fused into a single ossification, supporting the tail feathers and musculature. In modern birds, the rectrices attach to these. The pygostyle is the main component of the uropygium, a structure colloquially known as the bishop's nose, parson's nose, pope's nose, or sultan's nose. This is the fleshy protuberance visible at the posterior end of a bird (most commonly a chicken or turkey) that has been dressed for cooking. It has a swollen appearance because it also contains the uropygial gland that produces preen oil. Evolution Pygostyles probably began to evolve very early in the Cretaceous period, perhaps 140 – 130 million years ago. The earliest known species to have evolved a pygostyle were members of the Confuciusornithidae. The structure provided an evolutionary advantage, as a completely mobile tail as found in species such as '' Archaeopteryx'' is detrimental to its use for flight contro ...
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Omnivoropterygidae
Omnivoropterygidae (meaning "omnivorous wings") is a Family (biology), family of primitive avialans known exclusively from the Jiufotang Formation of China, though putative omnivoropterygids are known from the Maevarano Formation of the Maastrichtian of Madagascar. They had short skeletal tails and unusual skulls with teeth in the upper, but not lower, jaws. Their unique dentition has led some scientists to suggest an omnivore, omnivorous diet for them.Czerkas, S. A. & Ji, Q. (2002). "A preliminary report on an omnivorous volant bird from northeast China." ''In'': Czerkas, S. J. (editor): ''Feathered Dinosaurs and the origin of flight. The Dinosaur Museum Journal'' 1: 127-135HTML abstract/ref> The family was named by Stephen A. Czerkas & Qiang Ji in 2002, though its junior synonym Sapeornithidae is often used instead, though it was named four years later in 2006.Hu, D., Li, L., Hou, L., and Xu, X. (2010). "A new sapeornithid bird from China and its implication for early avian evolu ...
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Rachis
In biology, a rachis (from the grc, ῥάχις [], "backbone, spine") is a main axis or "shaft". In zoology and microbiology In vertebrates, ''rachis'' can refer to the series of articulated vertebrae, which encase the spinal cord. In this case the ''rachis'' usually forms the supporting axis of the body and is then called the spine or vertebral column. ''Rachis'' can also mean the central shaft of pennaceous feathers. In the gonad of the invertebrate nematode '' Caenorhabditis elegans'', a rachis is the central cell-free core or axis of the gonadal arm of both adult males and hermaphrodites where the germ cells have achieved pachytene and are attached to the walls of the gonadal tube. The rachis is filled with cytoplasm. In botany In plants, a rachis is the main axis of a compound structure. It can be the main stem of a compound leaf, such as in ''Acacia'' or ferns, or the main, flower-bearing portion of an inflorescence above a supporting peduncle. Where it subdivides ...
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