Crocodylomorphs
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Crocodylomorphs
Crocodylomorpha is a group of pseudosuchian archosaurs that includes the crocodilians and their extinct relatives. They were the only members of Pseudosuchia to survive the end-Triassic extinction. During Mesozoic and early Cenozoic times, crocodylomorphs were far more diverse than they are now. Triassic forms were small, lightly built, active terrestrial animals. The earliest and most primitive crocodylomorphs are represented by " sphenosuchians", a paraphyletic assemblage containing small-bodied forms with elongated limbs that walked upright, which represents the ancestral morphology of Crocodylomorpha. These forms persisted until the end of the Jurassic. During the Jurassic, Crocodylomorphs morphologically diversified into numerous niches, including into the aquatic and marine realms. Evolutionary history When their extinct species and stem group are examined, the crocodylian lineage (clade Pseudosuchia, formerly Crurotarsi) proves to have been a very diverse and adapti ...
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Carnufex
''Carnufex'' is an extinct genus of crocodylomorph suchian from the Late Triassic of North America. The genus was first described in 2015 by Zanno ''et al.'', who named the Binomial nomenclature, binomial ''Carnufex carolinensis'', meaning "Carolina butcher". Two specimens are known, the holotype skull and skeleton NCSM 21558, and the referred humerus NCSM 21623. The specimens are from the Carnian-age Pekin Formation, which dates to 231 million years ago. Based on the holotype, ''Carnufex'' would have been about long and tall, although it may have gotten larger due to the holotype not being fully grown. Discovery NCSM 21558 was discovered in a red Fluvial processes, fluvial Conglomerate (geology), conglomerate belonging to the mid-upper portion Pekin Formation of North Carolina, which formed in the Carnian age of the Late Triassic, around 231 million years ago. This specimen was described in a 2015 ''Scientific Reports'' article by Lindsay Zanno, Lindsay E. Zanno, Susan Drymal ...
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Terrestrisuchus
''Terrestrisuchus'' is an extinct genus of very small early crocodylomorph that was about long. Fossils have been found in Wales and Southern England and date from near the very end of the Late Triassic during the Rhaetian, and it is known by type and only known species ''T. gracilis''. ''Terrestrisuchus'' was a long-legged, active predator that lived entirely on land, unlike modern crocodilians. It inhabited a chain of tropical, low-lying islands that made up southern Britain, along with similarly small-sized dinosaurs and abundant rhynchocephalians. Numerous fossils of ''Terrestrisuchus'' are known from fissures in limestone karst which made up the islands it lived on, which formed caverns and sinkholes that preserved the remains of ''Terrestrisuchus'' and other island-living reptiles. Description ''Terrestrisuchus'' was a small, slender crocodylomorph with very long legs, quite unlike modern crocodilians. It was initially estimated to have been between long, although this e ...
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Saltoposuchus
''Saltoposuchus'' is an extinct genus of small (1-1.5 m and 10-15 kg), long-tailed crocodylomorph reptile (Sphenosuchia), from the Norian (Late Triassic) of Europe. The name translated means "leaping foot crocodile". It has been proposed that ''Terrestrisuchus gracilis'' and ''Saltoposuchus connectens'' represent different ontogenetic stages of the same genus. ''Saltoposuchus'' was commonly (and incorrectly) referred to in popular literature as the ancestor (or close ancestors) to dinosaurs, however, recent scientific research show that this is not the case. Description and Paleobiology Fossil evidence of Sphenosuchia and early crocodylomorphs lead paleontologists to conclude that ''Saltoposuchus'' is a terrestrial animal. As a monophyletic group of crocodylomorphs, Saltoposuchidae have many key morphological traits shared with most crocodylomorphs. Skull Much like other crocodylomorphs, ''Saltoposuchus'' skulls had a (reduced) antorbital fenestra, an overhanging squa ...
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Thalattosuchia
Thalattosuchia is a clade of marine crocodylomorphs from the Early Jurassic to the Early Cretaceous that had a cosmopolitan distribution. They are colloquially referred to as marine crocodiles or sea crocodiles, though they are not members of Crocodilia and records from Thailand and China suggest that some members lived in freshwater. The clade contains two major subgroupings, the Teleosauroidea and Metriorhynchoidea. Teleosauroids are not greatly specialised for oceanic life, with back osteoderms similar to other crocodyliformes. Within Metriorhynchoidea, the Metriorhynchidae displayed extreme adaptions for life in the open ocean, including the transformation of limbs into flippers, the development of a tail fluke, and smooth, scaleless skin. Discovery and naming The term Thalattosuchia was coined by Fraas in 1901.Fraas E. 1901. Die Meerkrokodile (Thalattosuchia n. g.) eine neue Sauriergruppe der Juraformation. ''Jahreshefte des Vereins für vaterländische Naturkunde, Württem ...
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Litargosuchus
''Litargosuchus'' is a sphenosuchian Crocodylomorpha, crocodylomorph, a Basal (phylogenetics), basal member of the Crocodylomorpha, crocodylomorph clade from the Early Jurassic of South Africa. Its genus name ''Litargosuchus'' is derived from Greek meaning "fast running crocodile" and its species name ''leptorhynchus'' refers to its Gracility, gracile snout. ''Litargosuchus'', along with all of South Africa's Crocodylomorpha, crocodylomorph Taxon, taxa, are confined to the upper Elliot Formation (UEF) in South Africa. History of discovery In 1988 James Kitching found the holotype fossil of ''Litargosuchus'' in a field expedition in Eagles Crag, a farm near Barkly East in the Eastern Cape. The material recovered consisted of a near-complete skull with its mandible and several bones belonging to the postcranial skeleton. At the time, James Kitching, Kitching was the director of the Evolutionary Studies Institute (previously the Bernard Price Institute) of the University of the Witw ...
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Archosaur
Archosauria () is a clade of diapsids, with birds and crocodilians as the only living representatives. Archosaurs are broadly classified as reptiles, in the cladistic sense of the term which includes birds. Extinct archosaurs include non-avian dinosaurs, pterosaurs, and extinct relatives of crocodilians. Modern paleontologists define Archosauria as a crown group that includes the most recent common ancestor of living birds and crocodilians, and all of its descendants. The base of Archosauria splits into two clades: Pseudosuchia, which includes crocodilians and their extinct relatives, and Avemetatarsalia, which includes birds and their extinct relatives (such as non-avian dinosaurs and pterosaurs). Older definitions of the group Archosauria rely on shared morphological characteristics, such as an antorbital fenestra in the skull, serrated teeth, and an upright stance. Some extinct reptiles, such as proterosuchids and euparkeriids, possessed these features yet originated pri ...
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Syntomiprosopus
''Syntomiprosopus'' (meaning "short face") is an extinct genus of archosauriform, possibly a crocodylomorph from the Late Triassic period of Arizona. The type and only known species is ''S. sucherorum''. ''Syntomiprosopus'' was unusually short-snouted, comparable to the Late Cretaceous notosuchian ''Simosuchus'', and is regarded as an example of convergent evolution between Triassic stem-archosaurs and Cretaceous archosaurs. Discovery and naming Fossils of ''Syntomiprosopus'' were discovered at a locality in the Downs' Quarry located in eastern Arizona, just above the "''Placerias'' Quarry" and just below the principal horizon of the Downs' Quarry. The fossils were recovered during joint excavations by the North Carolina Museum of Natural Sciences (NCSM) and Appalachian State University (ASU) between 2013 and 2015 as part of a series of excavations that had begun in 2010. The quarry is part of the expansive Late Triassic Chinle Formation, and has been radiometrically dated to ...
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Triassic–Jurassic Extinction Event
The Triassic–Jurassic (Tr-J) extinction event, often called the end-Triassic extinction, marks the boundary between the Triassic and Jurassic periods, , and is one of the top five major extinction events of the Phanerozoic eon, profoundly affecting life on land and in the oceans. In the seas, the entire Class (biology), class of conodonts and 23–34% of marine genus, genera disappeared. On land, all Archosauromorpha, archosauromorphs other than Crocodylomorpha, crocodylomorphs, pterosaurs, and dinosaurs became extinct; some of the groups which died out were previously abundant, such as aetosaurs, phytosaurs, and Rauisuchidae, rauisuchids. Some remaining non-mammalian therapsids and many of the large Temnospondyli, temnospondyl amphibians had become extinct prior to the Jurassic as well. However, there is still much uncertainty regarding a connection between the Tr-J boundary and terrestrial vertebrates, due to a lack of terrestrial fossils from the Rhaetian (latest) stage of the ...
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Pseudosuchia
Pseudosuchia is one of two major divisions of Archosauria, including living crocodilians and all archosaurs more closely related to crocodilians than to birds. Pseudosuchians are also informally known as "crocodilian-line archosaurs". Prior to 2011, the clade Pseudosuchia was often called Crurotarsi in reference to the crurotarsal ankle found in almost all members of the group, which traditionally included phytosaurs, ornithosuchids, and suchians. However, a major 2011 study of Triassic archosaur relations proposed that phytosaurs were not closely related to other traditional "crurotarsans", at least compared to "bird-line archosaurs" (Avemetatarsalians) such as pterosaurs and dinosaurs. As a result, the possession of a crurotarsal ankle was considered a plesiomorphic ("primitive") feature retained by pseudosuchians. Crurotarsi now refers to a broader group of reptiles including Pseudosuchia, Phytosauria, and Avemetatarsalia. Despite Pseudosuchia meaning "false crocodiles", th ...
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Sphenosuchus
''Sphenosuchus'' ("wedge crocodile") is an extinct genus of crocodylomorph from the Early Jurassic Elliot Formation of South Africa, discovered and described early in the 20th century. The skull is preserved very well but other than elements of the forelimb and isolated parts of the hind limb, the ''Sphenosuchus'' material is incomplete. It was probably quadrupedal, but may have been a facultative biped. ''Sphenosuchus'' was first thoroughly described in 1972 by the British palaeontologist Alick Walker, in a paper in the journal ''Nature''. Walker suggested, based on detailed (but still preliminary at that time) studies of the skull of ''Sphenosuchus'' and modern birds, that crocodylomorphs and birds might share an immediate common ancestor. Walker recanted his hypothesis in 1985, but restated and elaborated on it (in essence 'de-recanting') in a monograph published in 1990, which provided the most comprehensive description and discussion of ''Sphenosuchus'' yet published. Bro ...
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Phyllodontosuchus
''Phyllodontosuchus'' (Ancient Greek, meaning "leaf tooth crocodile", in reference to the shape of the middle and posterior teeth) is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from a skull and jaws from Lower Jurassic rocks of Yunnan, China. ''Phyllodontosuchus'' is unusual because some of its teeth were leaf-shaped, like those of some herbivorous dinosaurs, and it does not appear to have been a strict carnivore like most other crocodylomorphs. History and description ''Phyllodontosuchus'' is based on BVP568-L12, a crushed skull and jaws recovered from the Sinemurian-age Dark Red Beds of the Lufeng Formation near Dawa in Yunnan. A '' Morganucodon'' skull was also recovered from this locality. BVP568-L12 is only long and imperfectly preserved; it was first thought to represent an early ornithischian dinosaur. Sutures are not visible, so despite the small size, the specimen appea ...
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Dibothrosuchus
''Dibothrosuchus'' is a genus of sphenosuchian, a type of basal crocodylomorph, the clade that comprises the crocodilians and their closest kin. It is known from several partial skeletons and skulls. These fossils were found in Lower Jurassic (Lower Pliensbachian, circa 186 million years old) rocks of Yunnan, China. ''Dibothrosuchus'' was a small terrestrial crocodylomorph that probably had a keen sense of hearing, and thus was probably a vocal animal like modern crocodilians. History and description ''Dibothrosuchus'' was named in 1965 by D.J. Simmons based on a partial skull and skeleton collected by Father Oehler of Fu Jen Catholic University from mudstones near Huangchiatien. The type species is ''D. elaphros''. Additional remains of this genus were recovered in 1985 by a joint Chinese-US expedition. Fossils of ''Dibothrosuchus'' come from the Zhangjiawa Formation, being originally assigned to the Reds Beds of the Lufeng Formation, thus being geologically young ...
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