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Conodonts
Conodonts (Greek ''kōnos'', "cone", + ''odont'', "tooth") are an extinct group of agnathan (jawless) vertebrates resembling eels, classified in the class Conodonta. For many years, they were known only from their tooth-like oral elements, which are usually found in isolation and are now called conodont elements. Knowledge about soft tissues remains limited. They existed in the world's oceans for over 300 million years, from the Cambrian to the beginning of the Jurassic. Conodont elements are widely used as index fossils, fossils used to define and identify geological periods. The animals are also called Conodontophora (conodont bearers) to avoid ambiguity. Discovery and understanding of conodonts The teeth-like fossils of the conodont were first discovered by Heinz Christian Pander and the results published in Saint Petersburg, Russia, in 1856. The name ''pander'' is commonly used in scientific names of conodonts. It was only in the early 1980s that the first fossil evidence of ...
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Conodont Biostratigraphy
Conodonts are an extinct class of animals whose feeding apparatuses called ''teeth'' or ''elements'' are common microfossils found in strata dating from the Stage 10 of the Furongian, the fourth and final series of the Cambrian, to the Rhaetian stage of the Late Triassic. These elements can be used alternatively to or in correlation with other types of fossils (graptolites, trilobites, ammonites, ...) in the subfield of the stratigraphy named biostratigraphy. Paleozoic conodonts Cambrian conodonts It is suggested that ''Eoconodontus notchpeakensis'' can be a marker of the Stage 10 of the Furongian, the fourth and final series of the Cambrian. In 2006, a working group proposed the first appearance of '' Cordylodus andresi''. Currently the first appearance of ''E. notchpeakensis'' is favored by many authors because it is globally widespread and is independent of facies (known from continental rise to peritidal environments). The ''Eoconodontus notchpeakensis'' proposal would a ...
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Geological Periods
The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochronology (scientific branch of geology that aims to determine the age of rocks). It is used primarily by Earth scientists (including geologists, paleontologists, geophysicists, geochemists, and paleoclimatologists) to describe the timing and relationships of events in geologic history. The time scale has been developed through the study of rock layers and the observation of their relationships and identifying features such as lithologies, paleomagnetic properties, and fossils. The definition of standardized international units of geologic time is the responsibility of the International Commission on Stratigraphy (ICS), a constituent body of the International Union of Geological Sciences (IUGS), whose primary objective is to precisely define ...
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Agnathans
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (cyclostomes) and extinct (conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfish plus vertebrates. While a few scientist ...
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Agnatha
Agnatha (, Ancient Greek 'without jaws') is an infraphylum of jawless fish in the phylum Chordata, subphylum Vertebrata, consisting of both present (cyclostomes) and extinct ( conodonts and ostracoderms) species. Among recent animals, cyclostomes are sister to all vertebrates with jaws, known as gnathostomes. Recent molecular data, both from rRNA and from mtDNA as well as embryological data, strongly supports the hypothesis that living agnathans, the cyclostomes, are monophyletic. The oldest fossil agnathans appeared in the Cambrian, and two groups still survive today: the lampreys and the hagfish, comprising about 120 species in total. Hagfish are considered members of the subphylum Vertebrata, because they secondarily lost vertebrae; before this event was inferred from molecular and developmental data, the group Craniata was created by Linnaeus (and is still sometimes used as a strictly morphological descriptor) to reference hagfish plus vertebrates. While a few scie ...
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Ozarkodinida
Ozarkodinida is an extinct conodont order. It is part of the clade Prioniodontida, also known as the "complex conodonts". Name Ozarkodinida is named after the Ozark Mountains of Missouri Missouri is a U.S. state, state in the Midwestern United States, Midwestern region of the United States. Ranking List of U.S. states and territories by area, 21st in land area, it is bordered by eight states (tied for the most with Tennessee ..., United States. Elements The feeding apparatus of ozarkodinids is composed at the front of an axial Sa element, flanked by two groups of four close-set elongate Sb and Sc elements which were inclined obliquely inwards and forwards. Above these elements lay a pair of arched and inward pointing (makellate) M elements. Behind the S-M array lay transversely oriented and bilaterally opposed (pectiniform, i.e. comb-shaped) Pb and Pa elements. References External links * * Prehistoric jawless fish orders {{Conodont-stub ...
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Promissum
''Promissum'' is an extinct genus of conodonts, primitive chordates, that lived during the Upper Ordovician period. A conodont, ''Promissum'' had a primitive mouth under its eyes with mineralized teeth, which are both typical for conodonts. It had a primitive backbone and probably looked like a small eel or large worm, lacking any kind of fins except for perhaps a small one on the tail. It was relatively large for a conodont, reaching about 40 cm (16 inch) in length. Well-preserved specimens were discovered in the Soom shale of South Africa in 1994. ''Promissum'' was probably capable of maintaining a cruising speed, but not of bursts of speed. References External links ''Promissum''at fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals .... org (retrieved 30 A ...
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Promissum NT Small
''Promissum'' is an extinct genus of conodonts, primitive chordates, that lived during the Upper Ordovician period. A conodont, ''Promissum'' had a primitive mouth under its eyes with mineralized teeth, which are both typical for conodonts. It had a primitive backbone and probably looked like a small eel or large worm, lacking any kind of fins except for perhaps a small one on the tail. It was relatively large for a conodont, reaching about 40 cm (16 inch) in length. Well-preserved specimens were discovered in the Soom shale of South Africa in 1994. ''Promissum'' was probably capable of maintaining a cruising speed, but not of bursts of speed. References External links ''Promissum''at fossilworks Fossilworks is a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database The Paleobiology Database is an online resource for information on the distribution and classification of fossil animals .... org (retrieved 30 A ...
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Heinz Christian Pander
Heinz Christian Pander, also Christian Heinrich Pander ( – ), was a Russian Empire ethnic Baltic German biologist and embryologist. Biography In 1817 he received his doctorate from the University of Würzburg, and spent several years (1827–1842), performing scientific research from his estate in Carnikava (german: Zarnikau) on the banks of the Gauja River near Riga. In 1820 he took part in a scientific expedition to Bokhara as a naturalist. In 1826 he became a member of the Saint Petersburg Academy of Sciences. Research Pander studied the chick embryo and discovered the germ layers (i.e., three distinct regions of the embryo that give rise to the specific organ system). Because of these findings, he is considered by many to be the "founder of embryology". His work in embryology was continued by Karl Ernst von Baer (1792-1876), who expanded Pander's concept of germ layers to include all vertebrates. Pander performed important studies in the field of paleontology, being k ...
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Hydroxylapatite
Hydroxyapatite, also called hydroxylapatite (HA), is a naturally occurring mineral form of calcium apatite with the formula Ca5(PO4)3(OH), but it is usually written Ca10(PO4)6(OH)2 to denote that the crystal unit cell comprises two entities. Hydroxyapatite is the hydroxyl endmember of the complex apatite group. The OH− ion can be replaced by fluoride, chloride or carbonate, producing fluorapatite or chlorapatite. It crystallizes in the hexagonal crystal system. Pure hydroxyapatite powder is white. Naturally occurring apatites can, however, also have brown, yellow, or green colorations, comparable to the discolorations of dental fluorosis. Up to 50% by volume and 70% by weight of human bone is a modified form of hydroxyapatite, known as bone mineral. Carbonated calcium-deficient hydroxyapatite is the main mineral of which dental enamel and dentin are composed. Hydroxyapatite crystals are also found in pathological calcifications such as those found in breast tumors, as w ...
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Lagerstätte
A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, and Agua Nueva formations, the Eocene Green River Formation, the Miocene Foulden Maar and Ashfall F ...
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Mineralized Tissues
Mineralized tissues are biological tissues that incorporate minerals into soft matrices. Typically these tissues form a protective shield or structural support. Bone, mollusc shells, deep sea sponge ''Euplectella'' species, radiolarians, diatoms, antler bone, tendon, cartilage, tooth enamel and dentin are some examples of mineralized tissues. These tissues have been finely tuned to enhance their mechanical capabilities over millions of years of evolution. Thus, mineralized tissues have been the subject of many studies since there is a lot to learn from nature as seen from the growing field of biomimetics. The remarkable structural organization and engineering properties makes these tissues desirable candidates for duplication by artificial means. Mineralized tissues inspire miniaturization, adaptability and multifunctionality. While natural materials are made up of a limited number of components, a larger variety of material chemistries can be used to simulate the same propertie ...
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Notochord
In anatomy, the notochord is a flexible rod which is similar in structure to the stiffer cartilage. If a species has a notochord at any stage of its life cycle (along with 4 other features), it is, by definition, a chordate. The notochord consists of inner, vacuolated cells covered by fibrous and elastic sheaths, lies along the anteroposterior axis (''front to back''), is usually closer to the dorsal than the ventral surface of the embryo, and is composed of cells derived from the mesoderm. The most commonly cited functions of the notochord are: as a midline tissue that provides directional signals to surrounding tissue during development, as a skeletal (structural) element, and as a vertebral precursor. In lancelets the notochord persists throughout life as the main structural support of the body. In tunicates the notochord is present only in the larval stage, being completely absent in the adult animal. In these invertebrate chordates, the notochord is not vacuolated. In all ...
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