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Colobognatha
Colobognatha is a clade (formally considered a subterclass) of helminthomorph millipedes containing four orders: Platydesmida, Polyzoniida, Siphonocryptida, and Siphonophorida. Description The Colobognatha are united by several shared traits (synapomorphies). Males have two pairs of simple, leg-like gonopods, consisting of the rear leg pair of body segment 7 and the foreword-most leg pair of segment 8. They possess tubular defensive glands that open on the sides of the body, and lack a palp-like structure on their gnathochilaria. Other traits found in colobognathans, but not exclusively, include a distinctly narrow head, lack of Tömösváry organs, and no more than two pairs of ocelli A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-l .... References {{Taxonbar, from=Q1698087 ...
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Polyzoniida
__NOTOC__ Polyzoniida is an order of millipedes in the Class_(biology)#Hierarchy_of_ranks_below_and_above_the_level_of_class, subterclass Colobognatha containing three families and at least 74 described species. Description Polyzoniidans have a somewhat domed dorsal surface with a flat ventral side. Their heads are small and cone-like, with few ocelli. They lack a dorsal groove and paranota (lateral extensions of each segment). Classification The order Polyzoniida contains three families: :Hirudisomatidae Silvestri, 1896 (6 genera, 20 species) :Polyzoniidae Newport, 1844 (6 genera, 22 species) :Siphonotidae Cook, 1895 (15 genera, 40 species) Distribution The family Hirudisomatidae occurs from Spain to the Himalayas in Eurasia, Japan, and in North America from southwest Canada to central Mexico. Polyzoniidae has a holarctic (northern hemisphere) distribution, occurring in the Northwestern United States, northwest and Northeastern United States, northeast United States, eastern C ...
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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Ocelli
A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-lensed "compound eye", and is not necessarily at all simple in the usual sense of the word. The structure of an animal's eye is determined by the environment in which it lives, and the behavioural tasks it must fulfill to survive. Arthropods differ widely in the habitats in which they live, as well as their visual requirements for finding food or conspecifics, and avoiding predators. Consequently, an enormous variety of eye types are found in arthropods. They possess a wide variety of novel solutions to overcome visual problems or limitations. Use of the term ''simple eye'' is flexible, and must be interpreted in proper context; for example, the eyes of humans and of other large animals such as most cephalopods, are ''camera eyes'' and ...
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Gnathochilarium
In arthropods, the maxillae (singular maxilla) are paired structures present on the head as mouthparts in members of the clade Mandibulata, used for tasting and manipulating food. Embryologically, the maxillae are derived from the 4th and 5th segment of the head and the maxillary palps; segmented appendages extending from the base of the maxilla represent the former leg of those respective segments. In most cases, two pairs of maxillae are present and in different arthropod groups the two pairs of maxillae have been variously modified. In crustaceans, the first pair are called maxillulae (singular maxillula). Modified coxae at the base of the pedipalps in spiders are also called "maxillae", although they are not homologous with mandibulate maxillae. Myriapoda Millipedes In millipedes, the second maxillae have been lost, reducing the mouthparts to only the first maxillae which have fused together to form a gnathochilarium, acting as a lower lip to the buccal cavity and the man ...
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Palp
Pedipalps (commonly shortened to palps or palpi) are the second pair of appendages of chelicerates – a group of arthropods including spiders, scorpions, horseshoe crabs, and sea spiders. The pedipalps are lateral to the chelicerae ("jaws") and anterior to the first pair of walking legs. Overview Pedipalps are composed of six segments or articles: the coxa, the trochanter, the femur, the short patella, the tibia, and the tarsus. In spiders, the coxae frequently have extensions called maxillae or gnathobases, which function as mouth parts with or without some contribution from the coxae of the anterior legs. The limbs themselves may be simple tactile organs outwardly resembling the legs, as in spiders, or chelate weapons ( pincers) of great size, as in scorpions. The pedipalps of Solifugae are covered in setae, but have not been studied in detail. Comparative studies of pedipalpal morphology may suggest that leg-like pedipalps are primitive in arachnids. At present, the on ...
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Gonopod (millipede)
Gonopods are specialized appendages of various arthropods used in reproduction or egg-laying. In males, they facilitate the transfer of sperm from male to female during mating, and thus are a type of intromittent organ. In crustaceans and millipedes, gonopods are modified walking or swimming legs. Gonopods may be highly decorated with elaborate structures which may play roles in sperm competition, and can be used to differentiate and identify closely related species. Gonopods generally occur in one or more pairs, as opposed to the single (un-paired) reproductive organs such as the aedeagus of insects or the penis of harvestmen. Insects In insects, gonopods are appendages of the genital segment that may be used in insemination, or that comprise the egg-laying apparatus. Crustaceans In male decapod crustaceans, gonopods are modified swimming appendages (pleopods). The anterior two pair of pleopods in males are modified for sperm transferring, with differing degree of morphologi ...
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Synapomorphies
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Millipedes
Millipedes are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derived from this feature. Each double-legged segment is a result of two single segments fused together. Most millipedes have very elongated cylindrical or flattened bodies with more than 20 segments, while pill millipedes are shorter and can roll into a tight ball. Although the name "millipede" derives from the Latin for "thousand feet", no species was known to have 1,000 or more until the discovery of '' Eumillipes persephone'', which can have over 1,300 legs. There are approximately 12,000 named species classified into 16 orders and around 140 families, making Diplopoda the largest class of myriapods, an arthropod group which also includes centipedes and other multi-legged creatures. Most millipedes are slow-moving detritivores, eating decaying leaves and other dead plant matter. Some eat fungi o ...
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Johann Friedrich Von Brandt
Johann Friedrich von Brandt (25 May 1802 – 15 July 1879) was a German-Russian natural history, naturalist, who worked mostly in Russia. Brandt was born in Jüterbog and educated at a Gymnasium (school), gymnasium in Wittenberg and the Humboldt University of Berlin, University of Berlin. In 1831 he emigrated to Russia, and soon was appointed director of the Zoological Museum of the St Petersburg Academy of Sciences. Brandt encouraged the collection of native animals, many of which were not represented in the museum. Many specimens began to arrive from the expeditions of Nikolai Alekseevich Severtzov, Severtzov, Nikolai Przhevalsky, Przhevalsky, Aleksandr Fyodorovich Middendorf, Middendorff, Leopold von Schrenck, Schrenck and Gustav Radde. He described several birds collected by Russian explorers off the Pacific Coast of North America, including Brandt's cormorant, red-legged kittiwake and spectacled eider. As a paleontologist, Brandt ranks among the best. He was also an entomo ...
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Helminthomorpha
Chilognatha is a subclass of the class Diplopoda, which includes the vast majority of extant millipedes, about 12,000 species. Taxonomy The classification of Chilognatha presented below is based on Shear, 2011, and Shear & Edgecombe, 2010 (extinct groups). Recent cladistic and molecular studies have challenged the traditional classification schemes above, and in particular the position of the orders Siphoniulida and Polyzoniida is not yet well established. The placement and positions of extinct groups (†) known only from fossils is tentative and not fully resolved. After each name is listed the author citation: the name of the person who coined the name or defined the group, even if not at the current rank. * Subclass Chilognatha Latrielle, 1802 ** Order † Zosterogrammida Wilson, 2005 (Chilognatha ''incertae sedis'') ** Infraclass Pentazonia Brandt, 1833 *** Order †Amynilyspedida Hoffman, 1969 *** Superorder Limacomorpha Pocock, 1894 **** Order Glomeridesmida Cook, 18 ...
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Subterclass
In biological classification, class ( la, classis) is a taxonomic rank, as well as a taxonomic unit, a taxon, in that rank. It is a group of related taxonomic orders. Other well-known ranks in descending order of size are life, domain, kingdom, phylum, order, family, genus, and species, with class fitting between phylum and order. History The class as a distinct rank of biological classification having its own distinctive name (and not just called a ''top-level genus'' ''(genus summum)'') was first introduced by the French botanist Joseph Pitton de Tournefort in his classification of plants that appeared in his ''Eléments de botanique'', 1694. Insofar as a general definition of a class is available, it has historically been conceived as embracing taxa that combine a distinct ''grade'' of organization—i.e. a 'level of complexity', measured in terms of how differentiated their organ systems are into distinct regions or sub-organs—with a distinct ''type'' of constructio ...
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