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Coleopsis
''Coleopsis archaica'' is an extinct species of stem-group beetle. It is the only member of the genus ''Coleopsis'' and family Coleopsidae. It is known from a single specimen from the Early Permian (Asselian) of western Germany, estimated to be about 297 million years old. It is currently the oldest known beetle. Taxonomy and systematics While originally interpreted as a member of the family Tshekardocoleidae, this interpretation has been revised and questioned, with the most recent analyses suggesting that it is best placed in its own family, and may be the sister group to all other beetles. The family name was originally published in 2016 as "Coleopsidae" but a later publication claimed that this spelling was erroneous and proposed the spelling "Coleopseidae";Kirejtshuk, A.G. Taxonomic Review of Fossil Coleopterous Families (Insecta, Coleoptera). Suborder Archostemata: Superfamilies Coleopseoidea and Cupedoidea. Geosciences 2020, 10, 73. https://doi.org/10.3390/geosciences1002 ...
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Tshekardocoleidae
Tshekardocoleidae is an extinct family of beetles, known from the Permian. They represent amongst the oldest beetles. Like other primitive beetles, they are thought to have been xylophagous. They first appeared during the Cisuralian, before becoming extinct at the beginning of the Guadalupian. A claimed Jurassic record is doubtful. The oldest known beetle, ''Coleopsis,'' was originally assigned to this family, but is now assigned to its own family Coleopsidae. Taxonomy upundescribed Tshekardocoleid larvae, Tshekarda site Kirejtshuk (2020) included the following genera in Tshekardocoleidae * †'' Avocoleus'' - Obora site (Permian Sakmarian ); Moravia, Czech Republic **†'' Avocoleus fractus'' (Type species) **†'' Avocoleus neglegens'' * †'' Boscoleus'' - Obora site (Permian Sakmarian ); Moravia, Czech Republic **†'' Boscoleus blandus'' (Type species) * †'' Eocoleus'' - Obora site (Permian Sakmarian ); Moravia, Czech Republic **†'' Eocoleus scaber'' (Type ...
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Stem-group
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gr ...
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Permian
The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the sauropsids ( reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these drier conditions, rose to dominance in place of their am ...
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Asselian
In the geologic timescale, the Asselian is the earliest geochronologic age or lowermost chronostratigraphic stage of the Permian. It is a subdivision of the Cisuralian Epoch or Series. The Asselian lasted between and million years ago (Ma). It was preceded by the Gzhelian (the latest or uppermost subdivision in the Carboniferous) and followed by the Sakmarian. Stratigraphy The Asselian Stage was introduced into scientific literature in 1954, when the Russian stratigrapher V.E. Ruzhenchev split it from the Artinskian. At that moment the Artinskian still encompassed most of the lower Permian – its current definitions are more restricted. The Asselian is named after the Assel River in the southern Ural Mountains of Kazakhstan and Bashkortostan. The base of the Asselian Stage is at the same time the base of the Cisuralian Series and the Permian System. It is defined as the place in the stratigraphic record where fossils of the conodont ''Streptognathodus isolatus'' first appear ...
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Germany
Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated between the Baltic and North seas to the north, and the Alps to the south; it covers an area of , with a population of almost 84 million within its 16 constituent states. Germany borders Denmark to the north, Poland and the Czech Republic to the east, Austria and Switzerland to the south, and France, Luxembourg, Belgium, and the Netherlands to the west. The nation's capital and most populous city is Berlin and its financial centre is Frankfurt; the largest urban area is the Ruhr. Various Germanic tribes have inhabited the northern parts of modern Germany since classical antiquity. A region named Germania was documented before AD 100. In 962, the Kingdom of Germany formed the bulk of the Holy Roman Empire. During the 16th ce ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxonomic ...
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ICZN
The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the International Commission on Zoological Nomenclature (which shares the acronym "ICZN"). The rules principally regulate: * How names are correctly established in the frame of binominal nomenclature * Which name must be used in case of name conflicts * How scientific literature must cite names Zoological nomenclature is independent of other systems of nomenclature, for example botanical nomenclature. This implies that animals can have the same generic names as plants (e.g. there is a genus ''Abronia'' in both animals and plants). The rules and recommendations have one fundamental aim: to provide the maximum universality and continuity in the naming of all animals, except where taxonomic judgment dictates otherwise. The code is meant to guide o ...
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Tshekardocoleoidea
Tshekardocoleoidea is a superfamily in the extinct suborder Protocoleoptera that contains the following families: *† Coleopsidae *† Labradorocoleoidae *† Oborocoleidae *†Tshekardocoleidae Tshekardocoleidae is an extinct family of beetles, known from the Permian. They represent amongst the oldest beetles. Like other primitive beetles, they are thought to have been xylophagous. They first appeared during the Cisuralian, before becomi ... References Beetle superfamilies Taxa named by Boris Rohdendorf {{paleo-beetle-stub ...
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