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Coccotrema Maritimum
''Coccotrema maritimum'' is a crustose lichen commonly known as volcano lichen, due to the volcano-like appearance of the fruit bodies. It was first described and named by renowned Canadian lichenologist Irwin Brodo from a population on Haida Gwaii (for a brief time known as the Queen Charlotte Islands) in 1973. Description The thallus (the main body of the lichen) is thick, rough and crusty, taking on the appearance of dry, cracked mud. It is white to pinkish or yellowish-gray in colour, and porous to allow for gas exchange. From a distance, ''C. maritimum'' can be seen on the rocks as a white line just above the surface of the water. The outer surface of ''C.maritimum'' is scattered with hemispherical nitrogen-fixing structures (cephalodia) that are pinkish to brownish–grey in colour. Reproduction As in other lichens, ''Coccotrema'' species can reproduce sexually by generating tiny spores. Six to eight spores can be found per ascus (a sac where spores develop). Asci are ...
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Irwin Murray Brodo
Irwin M. Brodo (born 1935) is an emeritus scientist at the Canadian Museum of Nature, in Ottawa, Ontario, Canada. He is an authority on the identification and biology of lichens. Irwin Brodo was honored in 1994 with an Acharius Medal presented to him by the International Association for Lichenology. Brodo did his undergraduate studies at Columbia University, and received a master's degree from Cornell University. He earned a Ph.D. in lichenology under the supervision of Henry Imshaug at Michigan State University. He later went on to teach at Université Laval and the University of Alaska, and he also supervised master's students at the University of Ottawa and Carleton University. Brodo's list of publications includes 75 research papers, 8 popular articles, 22 reviews and 6 editorials and obituaries. In 1993, Brodie was awarded the Mary E. Elliot Service Award for his meritorious service to the Canadian Botanical Association. One of Irwin Brodo's great achievements was the pub ...
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Paraphyses
Paraphyses are erect sterile filament-like support structures occurring among the reproductive apparatuses of fungi, ferns, bryophytes and some thallophytes. The singular form of the word is paraphysis. In certain fungi, they are part of the fertile spore-bearing layer. More specifically, paraphyses are sterile filamentous hyphal end cells composing part of the hymenium of Ascomycota and Basidiomycota interspersed among either the asci or basidia respectively, and not sufficiently differentiated to be called cystidia A cystidium (plural cystidia) is a relatively large cell found on the sporocarp of a basidiomycete (for example, on the surface of a mushroom gill), often between clusters of basidia. Since cystidia have highly varied and distinct shapes that ar ..., which are specialized, swollen, often protruding cells. The tips of paraphyses may contain the pigments which colour the hymenium. In ferns and mosses, they are filament-like structures that are found on sporangia ...
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Lichens Of Western Canada
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichens Described In 1973
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

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Lichen Species
A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.Introduction to Lichens – An Alliance between Kingdoms
. University of California Museum of Paleontology.
Lichens have properties different from those of their component organisms. They come in many colors, sizes, and forms and are sometimes plant-like, but are not s. They may have tiny, leafless branches (); flat leaf-like structures (

Pertusariales
The Pertusariales are an order of fungi in the class Lecanoromycetes. It contains the following families: Agyriaceae The Agyriaceae are a family of lichenized fungi in the order Pertusariales. It contains two genera: ''Agyrium ''Agyrium'' is a genus of saprophytic fungi in the family Agyriaceae. It probably evolved from a lichen ancestor, as it is clos ..., Coccotremataceae, Icmadophilaceae, Megasporaceae, Microcaliciaceae, Ochrolechiaceae, Pertusariaceae, Varicellariaceae, and Variolariaceae. Many of these fungi form lichens. Gallery Image:Pertusaria_paratuberculifera_(EU).jpg, '' Pertusaria paratuberculifera'' (2 verrucae) Image:Pertusaria_paratuberculifera_(EU1).jpg, '' Pertusaria paratuberculifera'' (8 spores per ascus) References Lichen orders Lecanoromycetes orders Taxa named by David Leslie Hawksworth Taxa named by Maurice Choisy Taxa described in 1986 {{Pertusariales-stub ...
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Queen Charlotte Islands
Haida Gwaii (; hai, X̱aaydag̱a Gwaay.yaay / , literally "Islands of the Haida people") is an archipelago located between off the northern Pacific coast of Canada. The islands are separated from the mainland to the east by the shallow Hecate Strait. Queen Charlotte Sound lies to the south, with Vancouver Island beyond. To the north, the disputed Dixon Entrance separates Haida Gwaii from the Alexander Archipelago in the U.S. state of Alaska. Haida Gwaii consists of two main islands: Graham Island () in the north and Moresby Island (, literally: south people island half, or "Islands of Beauty") in the south, along with approximately 400 smaller islands with a total landmass of . Other major islands include Anthony Island ( / ), Burnaby Island (), Lyell Island, Louise Island, Alder Island ( / ), and Kunghit Island. (For a fuller, but still incomplete, list see List of islands of British Columbia.) Part of the Canadian province of British Columbia, the islands were known ...
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Ascus
An ascus (; ) is the sexual spore-bearing cell produced in ascomycete fungi. Each ascus usually contains eight ascospores (or octad), produced by meiosis followed, in most species, by a mitotic cell division. However, asci in some genera or species can occur in numbers of one (e.g. ''Monosporascus cannonballus''), two, four, or multiples of four. In a few cases, the ascospores can bud off conidia that may fill the asci (e.g. ''Tympanis'') with hundreds of conidia, or the ascospores may fragment, e.g. some ''Cordyceps'', also filling the asci with smaller cells. Ascospores are nonmotile, usually single celled, but not infrequently may be coenocytic (lacking a septum), and in some cases coenocytic in multiple planes. Mitotic divisions within the developing spores populate each resulting cell in septate ascospores with nuclei. The term ocular chamber, or oculus, refers to the epiplasm (the portion of cytoplasm not used in ascospore formation) that is surrounded by the "bourrelet ...
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Crustose
Crustose is a habit of some types of algae and lichens in which the organism grows tightly appressed to a substrate, forming a biological layer. ''Crustose'' adheres very closely to the substrates at all points. ''Crustose'' is found on rocks and tree bark. Some species of marine algae of the Rhodophyta, in particular members of the order Corallinales, family Corallinaceae, subfamily Melobesioideae with cell walls containing calcium carbonate grow to great depths in the intertidal zone, forming crusts on various substrates. The substrate can be rocks throughout the intertidal zone, or, as in the case of the Corallinales, reef-building corals, and other living organisms including plants, such as mangroves and animals such as shelled molluscs. The coralline red algae are major members of coral reef communities, cementing the corals together with their crusts. Among the brown algae, the order Ralfsiales comprises two families of crustose algae. Growth and habitat Many lichens gr ...
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Cephalodia
{{Short description, Morphological structure found in some lichens Cephalodia (singular cephalodium) are small gall-like structures found in some species of lichens that contain cyanobacterial symbionts Symbiosis (from Greek , , "living together", from , , "together", and , bíōsis, "living") is any type of a close and long-term biological interaction between two different biological organisms, be it mutualistic, commensalistic, or parasit .... Cephalodia can occur within the tissues of the lichen, or on its upper or lower surface. Lichens that have both green algal and cyanobacterial symbionts restrict the cyanobacteria to cephalodia. Lichens that have only cyanobacterial symbionts do not have cephalodia: instead the cyanobacteria are widely distributed throughout the lichen. Lichens with cephalodia can fix nitrogen, and can be an important contributor of nitrogen to the ecosystem. References *Brodo, I. M., S. D. Sharnoff, and S. Sharnoff. 2001. Lichens of North Ame ...
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Thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms were previously known as the thallophytes, a polyphyletic group of distantly related organisms. An organism or structure resembling a thallus is called thalloid, thallodal, thalliform, thalline, or thallose. A thallus usually names the entire body of a multicellular non-moving organism in which there is no organization of the tissues into organs. Even though thalli do not have organized and distinct parts (leaves, roots, and stems) as do the vascular plants, they may have analogous structures that resemble their vascular "equivalents". The analogous structures have similar function or macroscopic structure, but different microscopic structure; for example, no thallus has vascular tissue. In exceptional cases such as the Lemnoideae, where ...
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