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Clubmoss
Lycopodiopsida is a class of vascular plants also known as lycopsids, lycopods, or lycophytes. Members of the class are also called clubmosses, firmosses, spikemosses and quillworts. They have dichotomously branching stems bearing simple leaves called microphylls and reproduce by means of spores borne in sporangia on the sides of the stems at the bases of the leaves. Although living species are small, during the Carboniferous, extinct tree-like forms ( Lepidodendrales) formed huge forests that dominated the landscape and contributed to coal deposits. The nomenclature and classification of plants with microphylls varies substantially among authors. A consensus classification for extant (living) species was produced in 2016 by the Pteridophyte Phylogeny Group (PPG I), which places them all in the class Lycopodiopsida, which includes the classes Isoetopsida and Selaginellopsida used in other systems. (See Table 2.) Alternative classification systems have used ranks from div ...
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Euphyllophyte
The euphyllophytes are a clade of plants within the tracheophytes (the vascular plants). The group may be treated as an unranked clade, a division under the name Euphyllophyta or a subdivision under the name Euphyllophytina. The euphyllophytes are characterized by the possession of true leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, f ... ("megaphylls"), and comprise one of two major lineages of extant vascular plants. As shown in the cladogram below, the euphyllophytes have a sister relationship to the lycopodiophytes or lycopsids. Unlike the lycopodiophytes, which consist of relatively few presently living or extant taxa, the euphyllophytes comprise the vast majority of vascular plant lineages that have evolved since both groups shared a common ancestor more than 400 mill ...
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Lycopodiales
The Lycopodiaceae (class Lycopodiopsida, order Lycopodiales) are an old family of vascular plants, including all of the core clubmosses and firmosses, comprising 17 accepted genera and about 500 known species. This family originated about 380 million years ago in the early Devonian, though the diversity within the family has been much more recent. "Wolf foot" is another common name for this family due to the resemblance of either the roots or branch tips to a wolf's paw. Description Members of Lycopodiaceae are not spermatophytes and so do not produce seeds. Instead they produce spores, which are oily and flammable, and are the most economically important aspects of these plants. The spores are of one size (i.e. the plants are isosporous) and are borne on a specialized structure at the apex of a shoot called a strobilus (plural: strobili), which resembles a tiny battle club, from which the common name derives. Members of the family share the common feature of having a microphy ...
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Palhinhaea Cernua
''Palhinhaea cernua'', synonym ''Lycopodiella cernua'' and ''Lycopodium cernuum'', is a plant in the family Lycopodiaceae, commonly known as the staghorn clubmoss. The Hawaiian name for the plant is ''wāwaeʻiole'', or "rat's foot". It has a substantial number of scientific synonyms in several genera. The genus '' Palhinhaea'' is accepted in the Pteridophyte Phylogeny Group classification of 2016 (PPG I), but not in other classifications which submerge the genus in '' Lycopodiella''. It is the largest of the clubmosses, having rhizomes up to in length, with leafy uprights up to in height. Distribution ''Palhinhaea cernua'' is a widespread pan-tropical species, found mostly at higher elevations in subtropical mountain climates of tropical Africa, Asia, the Pacific Islands and the Neotropics. In Europe, it is found in the Azores (where it is possibly native) and formerly on Madeira. It has been introduced in continental Portugal (Valongo), Sicily and Malta Malta, offici ...
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Isoetes
''Isoetes'', commonly known as the quillworts, is a genus of lycopod. It is the only living genus in the family Isoetaceae and order Isoetales. , there were about 200 recognized species, with a cosmopolitan distribution mostly in aquatic habitats but with the individual species often scarce to rare. Species virtually identical to modern quillworts have existed since the Jurassic epoch, though the timing of the origin of modern ''Isoetes'' is subject to considerable uncertainty. The name of the genus may also be spelled ''Isoëtes''. The diaeresis (two dots over the e) indicates that the o and the e are to be pronounced in two distinct syllables. Including this in print is optional; either spelling (''Isoetes'' or ''Isoëtes'') is correct. Description Quillworts are mostly aquatic or semi-aquatic in clear ponds and slow-moving streams, though several (e.g. '' I. butleri'', '' I. histrix'' and '' I. nuttallii'') grow on wet ground that dries out in the summer. The ...
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Incertae Sedis
or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty at specific taxonomic levels is indicated by (of uncertain family), (of uncertain suborder), (of uncertain order) and similar terms. Examples * The fossil plant ''Paradinandra, Paradinandra suecica'' could not be assigned to any family, but was placed ''incertae sedis'' within the order Ericales when described in 2001. * The fossil ''Gluteus minimus (fossil), Gluteus minimus'', described in 1975, could not be assigned to any known animal phylum. The genus is therefore ''incertae sedis'' within the kingdom Animalia. * While it was unclear to which order the New World vultures (family Cathartidae) should be assigned, they were placed in Aves ''incertae sedis''. It was later agreed to place them in a separate order, Cathartiformes. * Boc ...
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Vascular Plant
Vascular plants (), also called tracheophytes (, ) or collectively tracheophyta (; ), are plants that have lignin, lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified Tissue (biology), tissue (the phloem) to conduct products of photosynthesis. The group includes most embryophyte, land plants ( accepted known species) excluding mosses. Vascular plants include the clubmosses, Equisetum, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). They are contrasted with nonvascular plants such as mosses and green algae. Scientific names for the vascular plants group include Tracheophyta, Tracheobionta and Equisetopsida sensu lato, Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "hi ...
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Plesion
In phylogenetics, the crown group or crown assemblage is a collection of species composed of the living representatives of the collection, the most recent common ancestor of the collection, and all descendants of the most recent common ancestor. It is thus a way of defining a clade, a group consisting of a species and all its extant or extinct descendants. For example, Neornithes (birds) can be defined as a crown group, which includes the most recent common ancestor of all modern birds, and all of its extant or extinct descendants. The concept was developed by Willi Hennig, the formulator of phylogenetic systematics, as a way of classifying living organisms relative to their extinct relatives in his "Die Stammesgeschichte der Insekten", and the "crown" and "stem" group terminology was coined by R. P. S. Jefferies in 1979. Though formulated in the 1970s, the term was not commonly used until its reintroduction in 2000 by Graham Budd and Sören Jensen. Contents of the crown gro ...
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Paraphyly
Paraphyly is a taxonomic term describing a grouping that consists of the grouping's last common ancestor and some but not all of its descendant lineages. The grouping is said to be paraphyletic ''with respect to'' the excluded subgroups. In contrast, a monophyletic grouping (a clade) includes a common ancestor and ''all'' of its descendants. The terms are commonly used in phylogenetics (a subfield of biology) and in the tree model of historical linguistics. Paraphyletic groups are identified by a combination of synapomorphies and symplesiomorphies. If many subgroups are missing from the named group, it is said to be polyparaphyletic. The term received currency during the debates of the 1960s and 1970s accompanying the rise of cladistics, having been coined by zoologist Willi Hennig to apply to well-known taxa like Reptilia (reptiles), which is paraphyletic with respect to birds. Reptilia contains the last common ancestor of reptiles and all descendants of that ancest ...
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Zosterophyll
The zosterophylls are a group of extinct land plants that first appeared in the Silurian period. The taxon was first established by Banks in 1968 as the subdivision Zosterophyllophytina; they have since also been treated as the division Zosterophyllophyta or Zosterophyta and the class or plesion Zosterophyllopsida or Zosteropsida. They were among the first vascular plants in the fossil record, and had a world-wide distribution. They were probably stem-group lycophytes, forming a sister group to the ancestors of the living lycophytes. By the late Silurian (late Ludlovian, about ) a diverse assemblage of species existed, examples of which have been found fossilised in what is now Bathurst Island in Arctic Canada. Morphology The stems of zosterophylls were either smooth or covered with small spines known as enations, branched dichotomously, and grew at the ends by unrolling, a process known as circinate vernation. The stems had a central vascular column in which the protox ...
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Bryophyte
Bryophytes () are a group of embryophyte, land plants (embryophytes), sometimes treated as a taxonomic Division (taxonomy), division referred to as Bryophyta ''Sensu#Common qualifiers, sensu lato'', that contains three groups of non-vascular plant, non-vascular land plants: the Marchantiophyta, liverworts, hornworts, and mosses. In the Sensu#Common qualifiers, strict sense, the division Bryophyta consists of the mosses only. Bryophytes are characteristically limited in size and prefer moist habitats although some species can survive in drier environments. The bryophytes consist of about 20,000 plant species. Bryophytes produce enclosed reproductive structures (gametangia and sporangia), but they do not produce flowers or seeds. They reproduce sexually by spores and asexually by fragmentation or the production of Gemma (botany), gemmae. Though bryophytes were considered a paraphyletic group in recent years, almost all of the most recent phylogenetics, phylogenetic evidence support ...
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Sporophyte
A sporophyte () is one of the two alternation of generations, alternating multicellular organism, multicellular phases in the biological life cycle, life cycles of plants and algae. It is a diploid multicellular organism which produces asexual Spore, spores. This stage Alternation of generations, alternates with a multicellular haploid gametophyte phase. Life cycle The sporophyte develops from the zygote produced when a haploid egg cell is fertilized by a haploid sperm and each sporophyte cell therefore has a double set of chromosomes, one set from each parent. All Embryophyta, land plants, and most multicellular algae, have life cycles in which a multicellular diploid sporophyte phase alternates with a multicellular haploid gametophyte phase. In the Spermatophyte, seed plants, the largest groups of which are the gymnosperms (bare seeds) and angiosperms (fruiting plants), the sporophyte phase is more prominent than the gametophyte, and is the familiar green plant with its roots, ...
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Myco-heterotrophy
Myco-heterotrophy (from Greek , , and ) is a symbiotic relationship between certain kinds of plants and fungi, in which the plant gets all or part of its food from parasitism upon fungi rather than from photosynthesis. A myco-heterotroph is the parasitic plant partner in this relationship. Myco-heterotrophy is considered a kind of cheating relationship and myco-heterotrophs are sometimes informally referred to as "mycorrhizal cheaters". This relationship is sometimes referred to as mycotrophy, though this term is also used for plants that engage in mutualistic mycorrhizal relationships. Relationship between myco-heterotrophs and host fungi Full (or obligate) myco-heterotrophy exists when a non-photosynthetic plant (a plant largely lacking in chlorophyll or otherwise lacking a functional photosystem) gets all of its food from the fungi that it parasitizes. Partial (or facultative) myco-heterotrophy exists when a plant is capable of photosynthesis, but parasitizes fung ...
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