Chamaesipho Grebneffi
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Chamaesipho Grebneffi
''Chamaesipho grebneffi'' is the first extinct member of the Notochthamalinae to be described, and the oldest chthamaloid barnacle known. This species is a fully derived ''Chamaesipho'', and indicates a considerable antiquity for Chthamaloidea. Three extinct species of '' Chthamalus'' from the middle Miocene of the faluns of Touraine, France were described in 2008 by Carriol. the New Zealand species is Oligocene, and considerably older. Discussion Like the three living species of ''Chamaesipho'', the shell of ''C. grebneffi'' begins with 6 plates, fusing completely at about 2 mm diameter. Freegrown specimens show a stellate basal outline. The scutum is highly distinctive in tergal margin equal in length to basal margin. In all other ''Chamaesipho'', tergal margin is much shorter than basal. The tergum is not definitely identified. ''Chamaesipho grebneffi'' is interpreted, from wear patterns on the shell, to have occupied high littoral The littoral zone or nearshore i ...
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Notochthamalinae
In the past, Notochthamalinae has been considered a subfamily of barnacles of the family Chthamalidae The Chthamalidae are a family of chthamaloid barnacles, living entirely in intertidal/subtidal habitats, characterized by a primary shell wall of eight, six, or four plates, lacking imbricating plate whorls, and either membraneous or more rare .... Research published in 2021 by Chan et al. resulted in the genera of the three subfamilies Euraphiinae, Chthamalinae, and Notochthamalinae being assigned directly to the family, without subfamily groupings, and this subfamily is no longer used. References {{Reflist, refs= {{Cite journal , title = The evolutionary diversity of barnacles, with an updated classification of fossil and living forms , date = 2021 , last1 = Chan , first1 = Benny K. K. , last2 = Dreyer , first2 = Niklas , last3 = Gale , first3 = Andy S. , last4 = Glenner , first4 = Henrik , last5 = Ewers-Saucedo , first5 = Christine , last6 = Pérez-Losada , ...
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Barnacle
A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosive settings. They are sessile (nonmobile) and most are suspension feeders, but those in infraclass Rhizocephala are highly specialized parasites on crustaceans. They have four nektonic (active swimming) larval stages. Around 1,000 barnacle species are currently known. The name is Latin, meaning "curl-footed". The study of barnacles is called cirripedology. Description Barnacles are encrusters, attaching themselves temporarily to a hard substrate or a symbiont such as a whale ( whale barnacles), a sea snake ('' Platylepas ophiophila''), or another crustacean, like a crab or a lobster (Rhizocephala). The most common among them, "acorn barnacles" ( Sessilia), are sessile where they grow their shells directly onto the substrate. Peduncul ...
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Chamaesipho
''Chamaesipho'' is a genus of four-plated notochthamaline barnacles in the Pacific Ocean limited to Australian/New Zealand temperate waters. They are intertidal in preference, and tend to form crowded columnar colonies. They can be identified in the field by having a four-plated wall, an unfused rostrum, and narrow opercular plates. ''Elminius'', which also inhabits the same area, has four plates in its shell wall. However, in ''Elminius'', the rostrum and rostrolatera are fused completely, and the compound rostrum receives the alae of the adjacent carinolaterals. In ''Chamaesipho'', the unfused rostrum bears alae, and closely resembles the carina in appearance. Definition and Discussion The primary shell wall is four plated, reduced from six by fusion of rostrolatera and carinolatera during ontogeny. No sutural interfolding is observed. With age, all plate sutures become concrescent. The basis is membraneous. Opercular plates are deeply articulated, but do not fuse together. Th ...
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Chthamaloidea
The Chthamaloidea are a subdivision of Balanomorpha proposed by Newman and Ross to include barnacles with shell wall composed of rostrum, carina, and one to three pairs of latera, rarely supplemented with one or more whorls of basal imbricating plates. The rostrolatus enters the sheath, but rarely fuses with the rostrum, as in the three higher superfamilies. Shell plates are simple in construction, solid, and incorporate organic chitin between carbonate layers. Opercular plates are deeply interlocked, and in some genera, may become concrescent with age. Soft part morphology includes concave labrum without notch in the central part. Cirrus III more resembles Cirrus IV than II, or may be intermediate in structure. Caudal appendages present in some species. Environmental preferences and distribution All living Chthamaloidea, and very sparsely known fossil occurrences inhabit surf zone/intertidal/subtidal littoral zone habitats. Geographically, chthamaloids may be found in all t ...
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Chthamalus
''Chthamalus'' (χθαμαλός, "flat" or "on the ground") is a genus of barnacles that is found along almost all non-boreal coasts of the northern hemisphere, as well as many regions in the southern hemisphere. These small barnacles have been studied in part because of the taxonomic confusion over a group of species that, by and large, are morphologically and ecologically quite similar. In recent years, molecular techniques have identified a number of cryptic species that have been subsequently confirmed by taxonomists using morphological measurements. Most recently the genus has been shown to be paraphyletic, with the genus ''Microeuraphia'' nested within ''Chthamalus''. Field Identification Identification of barnacles in the field can be challenging, and identification of particular species of ''Chthamalus'' is not considered reliable in the field other than a few species. However, a notable distinction in the arrangement of wall plates can help biologists distinguish Chth ...
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Miocene
The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" because it has 18% fewer modern marine invertebrates than the Pliocene has. The Miocene is preceded by the Oligocene and is followed by the Pliocene. As Earth went from the Oligocene through the Miocene and into the Pliocene, the climate slowly cooled towards a series of ice ages. The Miocene boundaries are not marked by a single distinct global event but consist rather of regionally defined boundaries between the warmer Oligocene and the cooler Pliocene Epoch. During the Early Miocene, the Arabian Peninsula collided with Eurasia, severing the connection between the Mediterranean and Indian Ocean, and allowing a faunal interchange to occur between Eurasia and Africa, including the dispersal of proboscideans into Eurasia. During the ...
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Oligocene
The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the exact dates of the start and end of the epoch are slightly uncertain. The name Oligocene was coined in 1854 by the German paleontologist Heinrich Ernst Beyrich from his studies of marine beds in Belgium and Germany. The name comes from the Ancient Greek (''olígos'', "few") and (''kainós'', "new"), and refers to the sparsity of extant forms of molluscs. The Oligocene is preceded by the Eocene Epoch and is followed by the Miocene Epoch. The Oligocene is the third and final epoch of the Paleogene Period. The Oligocene is often considered an important time of transition, a link between the archaic world of the tropical Eocene and the more modern ecosystems of the Miocene. Major changes during the Oligocene included a global expansion o ...
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Littoral
The littoral zone or nearshore is the part of a sea, lake, or river that is close to the shore. In coastal ecology, the littoral zone includes the intertidal zone extending from the high water mark (which is rarely inundated), to coastal areas that are permanently underwater, submerged — known as the ''foreshore'' — and the terms are often used interchangeably. However, the geographical meaning of ''littoral zone'' extends well beyond the intertidal zone to include all neritic waters within the bounds of continental shelves. Etymology The word ''littoral'' may be used both as a noun and as an adjective. It derives from the Latin language, Latin noun ''litus, litoris'', meaning "shore". (The doubled ''t'' is a late-medieval innovation, and the word is sometimes seen in the more classical-looking spelling ''litoral''.) Description The term has no single definition. What is regarded as the full extent of the littoral zone, and the way the littoral zone is divided into subregi ...
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