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Ceratomorpha
Odd-toed ungulates, mammals which constitute the taxonomic order Perissodactyla (, ), are animals—ungulates—who have reduced the weight-bearing toes to three (rhinoceroses and tapirs, with tapirs still using four toes on the front legs) or one (equines, third toe) of the five original toes. The non-weight-bearing toes are either present, absent, vestigial, or positioned posteriorly. By contrast, the even-toed ungulates bear most of their weight equally on four or two (an even number) of the five toes: their third and fourth toes. Another difference between the two is that odd-toed ungulates digest plant cellulose in their intestines rather than in one or more stomach chambers as even-toed ungulates, with the exception of Suina, do. The order includes about 17 species divided into three families: Equidae (horses, asses, and zebras), Rhinocerotidae (rhinoceroses), and Tapiridae (tapirs). Despite their very different appearances, they were recognized as related families i ...
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Chalicotheriidae
Chalicotheres (from Greek '' chalix'', "gravel" and '' therion'', "beast") are an extinct clade of herbivorous, odd-toed ungulate (perissodactyl) mammals that lived in North America, Eurasia, and Africa from the Middle Eocene until the Early Pleistocene, existing from 48.6 to 1.806 mya. They are one of the five major radiations of perissodactyls, with three groups living ( horses, plus the extinct paleotheres; rhinoceroses; tapirs), and two extinct (brontotheres and chalicotheres). Description Unlike modern perissodactyls, chalicotheres had clawed feet. They had longer forelimbs and shorter hind limbs, lower incisors that cropped food against a toothless pad in the upper jaw, low-crowned molar teeth, and were browsers on trees and shrubs throughout their history. They evolved in two different directions, which became separate subfamilies, the Schizotheriinae and the Chalicotheriinae. Schizotherine chalicotheres such as ''Moropus'' lived in a variety of forest, woodland, a ...
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Ungulate
Ungulates ( ) are members of the diverse clade Ungulata which primarily consists of large mammals with hooves. These include odd-toed ungulates such as horses, rhinoceroses, and tapirs; and even-toed ungulates such as cattle, pigs, giraffes, camels, sheep, deer, and hippopotamuses. Cetaceans such as whales, dolphins, and porpoises are also classified as even-toed ungulates, although they do not have hooves. Most terrestrial ungulates use the hoofed tips of their toes to support their body weight while standing or moving. The term means, roughly, "being hoofed" or "hoofed animal". As a descriptive term, "ungulate" normally excludes cetaceans as they do not possess most of the typical morphological characteristics of other ungulates, but recent discoveries indicate that they were also descended from early artiodactyls. Ungulates are typically herbivorous and many employ specialized gut bacteria to allow them to digest cellulose. Some modern species, such as pigs, are omnivorous, ...
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Tapiroidea
Tapiroidea is a superfamily of perissodactyls which includes the modern tapir. Members of the superfamily are small to large browsing mammals, roughly pig-like in shape, with short, prehensile snouts. Their closest relatives are the other odd-toed ungulates, including horses and rhinoceroses. Taxonomically, they are placed in suborder Ceratomorpha along with the rhino superfamily, Rhinocerotoidea. The first members of Tapiroidea appeared during the Early Eocene, 55 million years ago. Evolution and natural history The first tapirids, such as ''Heptodon'', appeared in the early Eocene. They appeared very similar to modern forms, but were about half the size, and lack the proboscis. The first true tapirs, appeared in the Oligocene, and by the Miocene, such genera as ''Miotapirus'' were almost indistinguishable to the extant species. It is believed that Asian and American tapirs diverged around 20 to 30 million years ago, and that tapirs migrated from North America and Central Ameri ...
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Rhinocerotoidea
Rhinocerotoidea is a superfamily (taxonomy), superfamily consisting of three family groups of odd-toed ungulates, three of which, the Amynodontidae, Hyracodontidae, and Paraceratheriidae, are extinct. The only extant family group is the Rhinocerotidae (true rhinoceroses), which survives as five living species. The extinct members of this superfamily are often called "rhinoceroses" alongside members of the family Rhinocerotidae, though they include genera, such as ''Paraceratherium'', which do not closely resemble modern rhinoceroses. Taxonomy The cladogram below follows a phylogenetic analysis by Bai ''et al.'' (2020): References

{{Taxonbar, from=Q15487229 Odd-toed ungulates Mammal superfamilies Taxa named by John Edward Gray ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Order (biology)
Order ( la, wikt:ordo#Latin, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between Family_(biology), family and Class_(biology), class. In biological classification, the order is a taxonomic rank used in the classification of organisms and recognized by the nomenclature codes. An immediately higher rank, superorder, is sometimes added directly above order, with suborder directly beneath order. An order can also be defined as a group of related families. What does and does not belong to each order is determined by a taxonomist, as is whether a particular order should be recognized at all. Often there is no exact agreement, with different taxonomists each taking a different position. There are no hard rules that a taxonomist needs to follow in describing or recognizing an order. Some taxa are accepted almost universally, while others are recognized only rarely. The name of an order is usually written with a capital letter. Fo ...
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Taxonomy (biology)
In biology, taxonomy () is the scientific study of naming, defining ( circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxa (singular: taxon) and these groups are given a taxonomic rank; groups of a given rank can be aggregated to form a more inclusive group of higher rank, thus creating a taxonomic hierarchy. The principal ranks in modern use are domain, kingdom, phylum (''division'' is sometimes used in botany in place of ''phylum''), class, order, family, genus, and species. The Swedish botanist Carl Linnaeus is regarded as the founder of the current system of taxonomy, as he developed a ranked system known as Linnaean taxonomy for categorizing organisms and binomial nomenclature for naming organisms. With advances in the theory, data and analytical technology of biological systematics, the Linnaean system has transformed into a system of modern biological classification intended to reflect the evolu ...
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Mammal
Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur or hair, and three middle ear bones. These characteristics distinguish them from reptiles (including birds) from which they diverged in the Carboniferous, over 300 million years ago. Around 6,400 extant species of mammals have been described divided into 29 orders. The largest orders, in terms of number of species, are the rodents, bats, and Eulipotyphla (hedgehogs, moles, shrews, and others). The next three are the Primates (including humans, apes, monkeys, and others), the Artiodactyla ( cetaceans and even-toed ungulates), and the Carnivora (cats, dogs, seals, and others). In terms of cladistics, which reflects evolutionary history, mammals are the only living members of the Synapsida (synapsids); this clade, together with Saur ...
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Rhinoceros Male 2003
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips to pl ...
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Rhinocerotidae
A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species of the superfamily Rhinocerotoidea.) Two of the extant species are native to Africa, and three to South and Southeast Asia. Rhinoceroses are some of the largest remaining megafauna: all weigh at least one tonne in adulthood. They have a herbivorous diet, small brains (400–600 g) for mammals of their size, one or two horns, and a thick (1.5–5 cm), protective skin formed from layers of collagen positioned in a lattice structure. They generally eat leafy material, although their ability to ferment food in their hindgut allows them to subsist on more fibrous plant matter when necessary. Unlike other perissodactyls, the two African species of rhinoceros lack teeth at the front of their mouths; they rely instead on their lips t ...
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Paraceratheriidae
Paraceratheriidae is an extinct family of long-limbed, hornless rhinocerotoids, commonly known as paraceratheres or indricotheres, that originated in the Eocene epoch and lived until the early Miocene. The first paraceratheres were only about the size of large dogs, growing progressively larger in the late Eocene and Oligocene. They were most common in the rainforest floodplain region which is now Kazakhstan, India, and southwest China, and lived further inland throughout northern and central Asia as well. The paraceratheres reached the peak of their evolution from the middle Oligocene to the early Miocene, where they became very large, herbivorous mammals. Most genera were about the size of modern draft horses and the extinct giant horse '' Equus giganteus'', with some growing significantly larger. The largest genus was ''Paraceratherium'', which was more than twice as heavy as a bull African elephant, and was one of the largest land mammals that ever lived. However, they remain ...
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Hyracodontidae
The Hyracodontidae are an extinct family of rhinoceroses endemic to North America, Europe, and Asia during the Eocene through early Oligocene, living from 48.6 to 26.3 million years ago (Mya), existing about . The Hyracodontidae thrived in the rainforests of Kazakhstan, Pakistan, and southwest China Southwest China () is a region in the south of the People's Republic of China. Geography Southwest China is a rugged and mountainous region, transitioning between the Tibetan Plateau to the west and the Chinese coastal hills (东南丘陵) and ..., a former coastal region. Fossil evidence also extends their geographical range to Germany. References Further reading * Lucas, S. G. & Sobus, J. C., (1989), The Systematics of Indricotheres. 358–378 in Prothero, D. R. & Schoch, R. M., (eds.) 1989: The Evolution of Perissodactyls, Oxford University Press, New York, New York & Oxford, England. Prehistoric rhinoceroses Oligocene odd-toed ungulates Rupelian extinctions Prehist ...
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