Cephalorhyncha Liticola
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Cephalorhyncha Liticola
Scalidophora is a group of marine pseudocoelomate protostomes that was proposed on morphological grounds to unite three phyla: the Kinorhyncha, the Priapulida and the Loricifera. The three phyla have four characters in common — chitinous cuticle that is moulted, rings of scalids on the introvert, flosculi, and two rings of introvert retracts.Heiner, I., Kristensen, R.H. 2005. Two new species of the genus ''Pliciloricus'' (Loricifera, Pliciloricidae) from the Faroe Bank, North Atlantic. Zoologischer Anzeiger. 243: 121–138. However, the monophyly of the Scalidophora was not supported by two molecular studies, where the position of the Loricifera was uncertain or as sister to the Panarthropoda. Both studies supported a reduced Scalidophora comprising the Kinorhyncha and Priapulida as sister phyla. Their closest relatives are the Panarthropoda, Nematoda and Nematomorpha; thus, they are placed in the group Ecdysozoa. The two species in the genus '' Markuelia'', known fro ...
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Ottoia Prolifica
''Ottoia'' is a stem-group archaeopriapulid worm known from Cambrian fossils. Although priapulid-like worms from various Cambrian deposits are often referred to ''Ottoia'' on spurious grounds, the only clear ''Ottoia'' macrofossils come from the Burgess Shale of British Columbia, which was deposited . Microfossils extend the record of ''Ottoia'' throughout the Western Canada Sedimentary Basin, from the mid- to late- Cambrian. A few fossil finds are also known from China. Morphology ''Ottoia'' specimens are on average 8 centimeters in length. Both length and width show variation with contraction; shorter specimens often being wider than longer ones. The characteristic proboscis of priapulids is present at the anterior, attached to the trunk of the animal, proceeded by the "bursa" at the posterior. The organism's body is bilaterally symmetrical, however, its anterior displays external radial symmetry. Like some other modern invertebrates, a cuticle restricts the size of and ...
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Cambrian
The Cambrian Period ( ; sometimes symbolized C with bar, Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Ca ...
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Markuelia
''Markuelia'' is a genus of fossil worm-like bilaterian animals allied to Ecdysozoa and known from strata of Lower Cambrian to Lower Ordovician age containing five species. An advanced X-ray imaging technique called X-ray tomographic microscopy has been applied to splendidly preserved, uncrushed ''Markuelia'' fossils found in Hunan province in southern China and in eastern Siberia. When details in features smaller than one micrometre across can be observed, these fossils are seen to represent many developmental stages, from the first cell divisions to the time of hatching; therefore they offer a unique opportunity to study the development of Lower Cambrian animals. The features observed indicate that the genus had a mouth surrounded by a ring of teeth, an alimentary canal, and an anus. However the characters preserved are inconclusive regarding the genus' taxonomic affinity; it can at best be placed in the scalidophoran total group, since it is currently impossible to ally ...
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Ecdysozoa
Ecdysozoa () is a group of protostome animals, including Arthropoda (insects, chelicerata, crustaceans, and myriapods), Nematoda, and several smaller phyla. They were first defined by Aguinaldo ''et al.'' in 1997, based mainly on phylogenetic trees constructed using 18S ribosomal RNA genes. A large study in 2008 by Dunn ''et al.'' strongly supported the Ecdysozoa as a clade, that is, a group consisting of a common ancestor and all its descendants. The group is also supported by morphological characters, and includes all animals that grow by ecdysis, moulting their cuticle. The group was initially contested by a significant minority of biologists. Some argued for groupings based on more traditional taxonomic techniques, while others contested the interpretation of the molecular data. Etymology The name ''Ecdysozoa'' stems etymologically . Characteristics The most notable characteristic shared by ecdysozoans is a three-layered cuticle (four in Tardigrada) composed of orga ...
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Nematomorpha
Nematomorpha (sometimes called Gordiacea, and commonly known as horsehair worms, hairsnakes, or Gordian worms) are a phylum of parasitoid animals superficially similar to nematode worms in morphology, hence the name. Most species range in size from long, reaching 2 metres in extreme cases, and in diameter. Horsehair worms can be discovered in damp areas, such as watering troughs, swimming pools, streams, puddles, and cisterns. The adult worms are free-living, but the larvae are parasitic on arthropods, such as beetles, cockroaches, mantises, orthopterans, and crustaceans. About 351 freshwater species are known and a conservative estimate suggests that there may be about 2000 freshwater species worldwide. The name "Gordian" stems from the legendary Gordian knot. This relates to the fact that nematomorphs often coil themselves in tight balls that resemble knots. Description and biology Nematomorphs possess an external cuticle without cilia. Internally, they have only longitudin ...
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Nematoda
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant-parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a broad range of environments. Less formally, they are categorized as Helminths, but are taxonomically classified along with arthropods, tardigrades and other moulting animals in the clade Ecdysozoa, and unlike flatworms, have tubular digestive systems with openings at both ends. Like tardigrades, they have a reduced number of Hox genes, but their sister phylum Nematomorpha has kept the ancestral protostome Hox genotype, which shows that the reduction has occurred within the nematode phylum. Nematode species can be difficult to distinguish from one another. Consequently, estimates of the number of nematode species described to date vary by author and may change rapidly over time. A 2013 survey of animal biodiversity published in the mega jou ...
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Panarthropoda
Panarthropoda is a proposed animal clade containing the extant phyla Arthropoda, Tardigrada (water bears) and Onychophora (velvet worms). Panarthropods also include extinct marine legged worms known as lobopodians ("Lobopodia"), a paraphyletic group where the last common ancestor and basal members (stem-group) of each extant panarthropod phylum are thought to have risen. However the term "Lobopodia" is sometimes expanded to include tardigrades and onychophorans as well. Common characteristics of the Panarthropoda include a segmented body, paired ladder-like, ventral nervous system, and the presence of paired appendages correlated with body segments. Taxonomy Not all studies support the monophyly of Panarthropoda, but most do, including neuroanatomical, phylogenomic and palaeontological studies. At least a close relationship between onychophorans and arthropods is widely agreed upon, but the position of tardigrades is more controversial. Some phylogenomic studies have f ...
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Philosophical Transactions Of The Royal Society B
''Philosophical Transactions of the Royal Society B: Biological Sciences'' is a biweekly peer-reviewed scientific journal published by the Royal Society. The editor-in-chief is John Pickett (Cardiff University). Overview Each issue covers a specific area of the biological sciences. Each issue aims to create an original and authoritative synthesis, often bridging traditional disciplines, which showcases current developments and provides a foundation for future research, applications and policy decisions. Each issue is edited by one or more expert guest editors. The themes fall into one of four general categories: * Cell and Development * Health and Disease * Environment and Evolution * Neuroscience and Cognition All articles become freely accessible one year after their publication date. History ''Philosophical Transactions of the Royal Society'' was established in 1665 by the Royal Society and is the oldest scientific journal in the English-speaking world. Henry Oldenburg ...
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Kinorhyncha
Kinorhyncha ( grc, κινέω, kīnéō, I move, ' "snout") is a phylum of small marine invertebrates that are widespread in mud or sand at all depths as part of the meiobenthos. They are also called mud dragons. Modern species are or less, but Cambrian forms could reach . Anatomy Kinorhynchs are limbless animals, with a body consisting of a head, neck, and a trunk of eleven segments. They are the only members of Ecdysozoa, except from the panarthropoda, with a segmented body. Juveniles have eight or nine segments, depending on genus, with the last two or three being added later during growth. Like other ecdysozoans they do not have external cilia, but instead have a number of spines along the body, plus up to seven circles of spines around the head. These spines are used for locomotion, withdrawing the head and pushing forward, then gripping the substrate with the spines while drawing up the body. The body wall consists of a thin syncitial layer, which secretes a tough ...
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Phylum (biology)
In biology, a phylum (; plural: phyla) is a level of classification or taxonomic rank below kingdom and above class. Traditionally, in botany the term division has been used instead of phylum, although the International Code of Nomenclature for algae, fungi, and plants accepts the terms as equivalent. Depending on definitions, the animal kingdom Animalia contains about 31 phyla, the plant kingdom Plantae contains about 14 phyla, and the fungus kingdom Fungi contains about 8 phyla. Current research in phylogenetics is uncovering the relationships between phyla, which are contained in larger clades, like Ecdysozoa and Embryophyta. General description The term phylum was coined in 1866 by Ernst Haeckel from the Greek (, "race, stock"), related to (, "tribe, clan"). Haeckel noted that species constantly evolved into new species that seemed to retain few consistent features among themselves and therefore few features that distinguished them as a group ("a self-contained uni ...
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Protostome
Protostomia () is the clade of animals once thought to be characterized by the formation of the organism's mouth before its anus during embryonic development. This nature has since been discovered to be extremely variable among Protostomia's members, although the reverse is typically true of its sister clade, Deuterostomia. Well known examples of protostomes are arthropods, molluscs, annelids, flatworms and nematodes. They are also called schizocoelomates since schizocoely typically occurs in them. Together with the Deuterostomia and Xenacoelomorpha, these form the clade Bilateria, animals with bilateral symmetry, anteroposterior axis and three germ layers. Protostomy In animals at least as complex as earthworms, the first phase in gut development involves the embryo forming a dent on one side (the blastopore) which deepens to become its digestive tube (the archenteron). In the sister-clade, the deuterostomes (), the original dent becomes the anus while the gut eventually tu ...
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