Camposaurus
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Camposaurus
''Camposaurus'' ( ) is a coelophysid dinosaur genus from the Norian stage of the Late Triassic period of North America. The pertinent fossil remains date back to the early to middle Norian stage, and is widely regarded as the oldest known neotheropod. Description ''Camposaurus'' is a small, carnivorous, theropod dinosaur. Its approximate length and weight cannot be reliably estimated because of the sparse material that is known from this genus. ''Camposaurus'' is known from partial lower leg bones, holotype UCMP 34498 (which includes distal tibiae, distal fibulae, and astragalocalcanea), and other fragmentary material. Like other coelophysids, it has fused tibio-tarsals and fibulo-tarsals. Unlike its relatives, the area of the tibia that fits with the fibula has a distinct ridge at the back. Another unique feature is the lack of a large medial condyle on the astragalus. The type species, ''C. arizonensis'', was formally named and described by Adrian Hunt, Spencer G. Lucas, Andre ...
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Camposaurus Arizonensis
''Camposaurus'' ( ) is a coelophysid dinosaur genus from the Norian stage of the Late Triassic period of North America. The pertinent fossil remains date back to the early to middle Norian stage, and is widely regarded as the oldest known neotheropod. Description ''Camposaurus'' is a small, carnivorous, theropod dinosaur. Its approximate length and weight cannot be reliably estimated because of the sparse material that is known from this genus. ''Camposaurus'' is known from partial lower leg bones, holotype UCMP 34498 (which includes distal tibiae, distal fibulae, and astragalocalcanea), and other fragmentary material. Like other coelophysids, it has fused tibio-tarsals and fibulo-tarsals. Unlike its relatives, the area of the tibia that fits with the fibula has a distinct ridge at the back. Another unique feature is the lack of a large medial condyle on the astragalus. The type species, ''C. arizonensis'', was formally named and described by Adrian Hunt, Spencer G. Lucas, An ...
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Coelophysid
Coelophysidae is a family of primitive carnivorous theropod dinosaurs. Most species were relatively small in size. The family flourished in the Late Triassic and Early Jurassic periods, and has been found on numerous continents. Many members of Coelophysidae are characterized by long, slender skulls and light skeletons built for speed. One member genus, ''Coelophysis'', displays the earliest known furcula in a dinosaur. Under cladistic analysis, Coelophysidae was first defined by Paul Sereno in 1998 as the most recent common ancestor of ''Coelophysis bauri'' and '' Procompsognathus triassicus'', and all of that common ancestor's descendants. However, Tykoski (2005) has advocated for the definition to change to include the additional taxa of "Syntarsus" ''kayentakatae'' and '' Segisaurus halli''. Coelophysidae is part of the superfamily Coelophysoidea, which in turn is a subset of the larger Neotheropoda clade. As part of Coelophysoidea, Coelophysidae is often placed as sister to ...
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Coelophysidae
Coelophysidae is a family of primitive carnivorous theropod dinosaurs. Most species were relatively small in size. The family flourished in the Late Triassic and Early Jurassic periods, and has been found on numerous continents. Many members of Coelophysidae are characterized by long, slender skulls and light skeletons built for speed. One member genus, ''Coelophysis'', displays the earliest known furcula in a dinosaur. Under Cladistics, cladistic analysis, Coelophysidae was first defined by Paul Sereno in 1998 as the most recent common ancestor of ''Coelophysis, Coelophysis bauri'' and ''Procompsognathus, Procompsognathus triassicus'', and all of that common ancestor's descendants. However, Tykoski (2005) has advocated for the definition to change to include the additional taxa of Coelophysis kayentakatae, "Syntarsus" ''kayentakatae'' and ''Segisaurus halli''. Coelophysidae is part of the superfamily Coelophysoidea, which in turn is a subset of the larger Neotheropoda clade. As p ...
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Coelophysis
''Coelophysis'' ( traditionally; or , as heard more commonly in recent decades) is an extinct genus of coelophysid theropod dinosaur that lived approximately 228 to 201.3 million years ago during the latter part of the Triassic Period from the Carnian and Rhaetian faunal stages in what is now the southwestern United States. ''Megapnosaurus'' was once considered a species within this genus,Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Jurassic, Africa)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 535–536. but this interpretation has been challenged since 2017 and the genus ''Megapnosaurus'' is now considered valid. ''Coelophysis'' was a small, slenderly-built, ground-dwelling, bipedal carnivore that could grow up to long. It is one of the earliest known dinosaur genera. Scattered material representing similar animals has been found worldwide in some Late Tria ...
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Coelophysis Rhodesiensis
''Megapnosaurus'' (meaning "big dead lizard", from Greek μεγα = "big", 'απνοος = "not breathing", "dead", σαυρος = "lizard") is an extinct genus of coelophysid theropod dinosaur that lived approximately 188 million years ago during the early part of the Jurassic Period in what is now Africa. The species was a small to medium-sized, lightly built, ground-dwelling, bipedal carnivore, that could grow up to long and weigh up to . It was originally given the genus name ''Syntarsus'', but that name was later determined to be preoccupied by a beetle. The species was subsequently given a new genus name, ''Megapnosaurus,'' by Ivie, Ślipiński & Węgrzynowicz in 2001. Some studies have classified it as a species within the genus ''Coelophysis,'' but this interpretation has been challenged by more subsequent studies and the genus is now considered valid. Discovery and history The first fossils of ''Megapnosaurus'' were found in 1963 by a group of students from Northlea S ...
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Bluewater Creek Formation
The Bluewater Creek Formation is a geologic formation in west-central New Mexico. It preserves fossils dating back to the late Triassic period. Description The formation consists of red sandstones and mudstones and has a total thickness in excess of . It conformably overlies the Shinarump Conglomerate and conformably underlies the Petrified Forest Formation. The formation has been correlated with the Salitral Formation of north-central New Mexico on the basis of its lithology, but it is likely somewhat younger, with an age around 219 million years. Fossils The formation contains some fossil vertebrates and an extensive fossil flora. Fossil tetrapods from the Bluewater Creek Formation include the theropod dinosaur ''Camposaurus'', phytosaur ''Rutiodon'', the aetosaurs ''Desmatosuchus'' and ''Stagonolepis'', and the metoposaurid ''Anaschisma''. These are characteristic of the late Carnian. History of investigation Spencer G. Lucas and S.N. Hayden defined the Bluewater Creek Mem ...
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Charles Lewis Camp
Charles Lewis Camp (March 12, 1893 Jamestown, North Dakota – August 14, 1975 San Jose, California) was a palaeontologist and zoologist, working from the University of California, Berkeley. He took part in excavations at the 'Placerias Quarry', in 1930 and the forty ''Shonisaurus'' skeleton discoveries of the 1960s, in what is now the Berlin-Ichthyosaur State Park. Camp served as the third director of the University of California Museum of Paleontology from 1930 to 1949, and coincidentally as chair of the UC Berkeley Paleontology Department between 1939 and 1949. Camp named a number of species of marine reptiles such as ''Shonisaurus'' and ''Plotosaurus'', as well as the dinosaur '' Segisaurus''. Camp was also an important bibliographer and historian of Western America. This aspect of his career is represented most notably by two works. The first is his biography of American pioneer James Clyman, which Bernard De Voto called "one of the half-dozen classics in the field." The secon ...
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Coelophysoids
Coelophysoidea were common dinosaurs of the Late Triassic and Early Jurassic periods. They were widespread geographically, probably living on all continents. Coelophysoids were all slender, carnivorous forms with a superficial similarity to the coelurosaurs, with which they were formerly classified, and some species had delicate cranial crests. Sizes range from about 1 to 6 m in length. It is unknown what kind of external covering coelophysoids had, and various artists have portrayed them as either scaly or feathered. Some species may have lived in packs, as inferred from sites where numerous individuals have been found together. Examples of coelophysoids include ''Coelophysis'', ''Procompsognathus'' and ''Liliensternus''. Most dinosaurs formerly referred to as being in the dubious taxon "Podokesauridae" are now classified as coelophysoids. Classification Despite their very early occurrence in the fossil record, coelophysoids have a number of derived features that separate the ...
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Nomen Dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium'' it may be impossible to determine whether a specimen belongs to that group or not. This may happen if the original type series (i. e. holotype, isotype, syntype or paratype) is lost or destroyed. The zoological and botanical codes allow for a new type specimen, or neotype, to be chosen in this case. A name may also be considered a ''nomen dubium'' if its name-bearing type is fragmentary or lacking important diagnostic features (this is often the case for species known only as fossils). To preserve stability of names, the ''International Code of Zoological Nomenclature'' allows a new type specimen, or neotype, to be chosen for a ''nomen dubium'' in this case. 75.5. Replacement of unidentifiable name-bearing type by a neotype. When an author considers that the taxonomic identity of a ...
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Autapomorphies
In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to the focal taxon (which may be a species, family or in general any clade). It can therefore be considered an apomorphy in relation to a single taxon. The word ''autapomorphy'', first introduced in 1950 by German entomologist Willi Hennig, is derived from the Greek words αὐτός, ''autos'' "self"; ἀπό, ''apo'' "away from"; and μορφή, ''morphḗ'' = "shape". Discussion Because autapomorphies are only present in a single taxon, they do not convey information about relationship. Therefore, autapomorphies are not useful to infer phylogenetic relationships. However, autapomorphy, like synapomorphy and plesiomorphy is a relative concept depending on the taxon in question. An autapomorphy at a given level may well be a synapomorphy at ...
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Ceratosauria
Ceratosaurs are members of the clade Ceratosauria, a group of dinosaurs defined as all theropods sharing a more recent common ancestor with ''Ceratosaurus'' than with birds. The oldest known ceratosaur, ''Saltriovenator'', dates to the earliest part of the Jurassic, around 199 million years ago. According to the majority of the latest research, Ceratosauria includes three major clades: Ceratosauridae, Noasauridae, and Abelisauridae, found primarily (though not exclusively) in the Southern Hemisphere. Originally, Ceratosauria included the above dinosaurs plus the Late Triassic to Early Jurassic Coelophysoidea and Dilophosauridae, implying a much earlier divergence of ceratosaurs from other theropods. However, most recent studies have shown that coelophysoids and dilophosaurids do not form a natural group with other ceratosaurs, and are excluded from this group. Ceratosauria derives its names from the type species, ''Ceratosaurus nasicornis'', described by O.C. Marsh in 1884. A ...
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Clade
A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English term, the equivalent Latin term ''cladus'' (plural ''cladi'') is often used in taxonomical literature. The common ancestor may be an individual, a population, or a species (extinct or extant). Clades are nested, one in another, as each branch in turn splits into smaller branches. These splits reflect evolutionary history as populations diverged and evolved independently. Clades are termed monophyletic (Greek: "one clan") groups. Over the last few decades, the cladistic approach has revolutionized biological classification and revealed surprising evolutionary relationships among organisms. Increasingly, taxonomists try to avoid naming taxa that are not clades; that is, taxa that are not monophyletic. Some of the relationships between organisms ...
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