Brachythoraci
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Brachythoraci
Brachythoraci is an extinct suborder of arthrodire placoderms, armored fish most diverse during the Devonian. Phylogeny Arthrodira is divided into three main groups: the paraphyletic Actinolepida and Phlyctaenii, and then the monophyletic Brachythoraci. Brachythoraci is then further divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Below is a cladogram from the 2016 Zhu ''et al.'' phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study: References Arthrodires Prehistoric animal suborders Fish suborders {{Placoderm-stub ...
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Arthrodire
Arthrodira (Greek for "jointed neck") is an Order (biology), order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus ...
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Phlyctaenioidei
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured ju ...
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Arthrodira
Arthrodira (Greek for "jointed neck") is an order of extinct armored, jawed fishes of the class Placodermi that flourished in the Devonian period before their sudden extinction, surviving for about 50 million years and penetrating most marine ecological niches. Arthrodires were the largest and most diverse of all groups of Placoderms. Description Arthrodire placoderms are notable for the movable joint between armor surrounding their heads and bodies. Like all placoderms, they lacked distinct teeth; instead, they used the sharpened edges of a bony plate on their jawbone as a biting surface. The eye sockets are protected by a bony ring, a feature shared by birds and some ichthyosaurs. Early arthrodires, such as the genus ''Arctolepis'', were well-armoured fishes with flattened bodies. The largest member of this group, ''Dunkleosteus'', was a true superpredator of the latest Devonian period, reaching as much as 6 m in length. In contrast, the long-nosed ''Rolfosteus'' measured just ...
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Eubrachythoraci
Eubrachythoraci is an extinct clade of arthrodire placoderms within the suborder Brachythoraci, armored fish most diverse during the Devonian. Most are considered to be pelagic (open ocean) long-distance swimmers, leading to their widespread distribution beginning from at least the Middle Devonian period. Phylogeny Brachythoraci is divided into the large derived clade Eubrachythoraci and several basal groups: Buchanosteoidea, Homosteidae, and Holonematidae. (Although Holonematidae's membership in Brachythoraci is disputed.) Eubrachythoraci is then further divided into the sub-clades Coccosteomorphi and Pachyosteomorphi, the latter of which can be further sub-divided into Aspinothoracidi and Dunkleosteoidea, as shown in the cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
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Coccosteina
Coccosteina is an extinct infraorder of placoderms, armored fish most diverse during the Devonian.Heintz., Anatol, 1932, The structure of ''Dinichthys'', a contribution to your knowledge of the Arthrodira: In: The Bashford Dean Memorial Volume Archaic Fishes'', edited by Gudger, E. W., article IV, The American Museum of Natural History, 224pp. However, the term is no longer in use, as modern cladistical methods have produced alternative phylogenetic trees of Brachythoraci with new subdivisions. Systematics * Basal genus '' Maideria'' * Basal genus ''Xiangshuiosteus'' * Superfamily Buchanosteoidea ** Family Buchanosteidae * Superfamily Gemuendenaspoidea ** Family Gemuendenaspidae * Superfamily Homosteoidea ** Family Homostiidae * Superfamily Brachydeiroidea ** Family Brachydeiridae ** Family Leptosteidae * Superfamily Coccosteoidea ** Family Pholidosteidae ** Family Coccosteidae ** Family Plourdosteidae ** Family Torosteidae ** Family Incisoscutidae ** Family Camuropisc ...
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Xiangshuiosteus
''Xiangshuiosteus wui'' is an extinct monospecific genus of brachythoracid arthrodire placoderm from the Late Emsian stage of the Early Devonian epoch, discovered in Wuding County of Yunnan province, China. It has recently been reassessed as a dunkleosteid. Etymology The generic name literally translates as "(Pinyin:Xiāngshuǐ) + bone," but actually refers to Xiangshui Valley,Not to be confused with Xiangshui County ()(Pinyin:Xiǎng shuǐ) in Jiangsu province the district in Wuding County where the holotype was found. The specific name honors Wu Baosheng, the gentleman who provided the holotype to Wang Junqing, the species' describer. Specimen and taxonomy ''X. wui'' is known from a flattened, "Buddhist cap" shaped skull roof. The skull roof is strongly reminiscent of those of coccosteids, but also has anatomical features otherwise diagnostic of buchanosteids. This mix of anatomy lead its describer, Wang Junqing, to suggest that ''X. wui'' is the sister taxon of Coccosteid ...
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Coccosteus Cuspidatus
''Coccosteus'' (from el, κόκκος , 'berry' and el, ὀστέον 'bone') is an extinct genus of arthrodire placoderm from the Devonian period. Its fossils have been found throughout Europe and North America. The majority of these have been found in freshwater sediments, though such a large range suggests that they may have been able to enter saltwater. The largest specimens were about , although the average length was . Description Like all other arthrodires, ''Coccosteus'' had a joint between the armor of the body and skull. It also had an internal joint between its neck vertebrae and the back of the skull, allowing for the mouth to be opened even wider. Along with the longer jaws, this allowed ''Coccosteus'' to feed on fairly large prey. The up-and-down movement of the skull also allowed for more water to be pumped through the gills. Possibly, the creature supplemented its diet with organic material filtered from mud using the gills. As with all other arthrodires, ''Cocc ...
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Homostiidae
Homostiidae is a family of flattened arthrodire placoderms from the Early to Middle Devonian. Fossils appear in various strata in Europe, Russia, Morocco, Australia, Canada and Greenland. Many homostiids have " toothless" jaws, and large sizes. suggesting that many homostiids were probably filter feeders, like the also noticeably flattened Rhincodon typus. All homostiids have flattened and elongated skulls. According to Denison 1978, primitive homostiids have moderately long median dorsal plates, whereas in "advanced" homostiids, the median dorsal tends to be short and broad. Obruchev (1964) placed the following primitive genera ''Euleptaspis'', ''Lophostracon'' and ''Luetkeichthys'' in a separate family, "Euleptaspididae," and Ørvig (1969), claimed that the Euleptaspidids were totally unrelated to Homostiidae proper (i.e., being neither related to, nor ancestral), but, according to Denison, did not clearly explain his reasons why this was so. Genera ''Angarichthys'' A compa ...
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Homosteidae
Homostiidae is a family of flattened arthrodire placoderms from the Early to Middle Devonian. Fossils appear in various strata in Europe, Russia, Morocco, Australia, Canada and Greenland. Many homostiids have " toothless" jaws, and large sizes. suggesting that many homostiids were probably filter feeders, like the also noticeably flattened Rhincodon typus. All homostiids have flattened and elongated skulls. According to Denison 1978, primitive homostiids have moderately long median dorsal plates, whereas in "advanced" homostiids, the median dorsal tends to be short and broad. Obruchev (1964) placed the following primitive genera ''Euleptaspis'', ''Lophostracon'' and ''Luetkeichthys'' in a separate family, "Euleptaspididae," and Ørvig (1969), claimed that the Euleptaspidids were totally unrelated to Homostiidae proper (i.e., being neither related to, nor ancestral), but, according to Denison, did not clearly explain his reasons why this was so. Genera ''Angarichthys'' A compa ...
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Basal (phylogeny)
In phylogenetics, basal is the direction of the ''base'' (or root) of a rooted phylogenetic tree or cladogram. The term may be more strictly applied only to nodes adjacent to the root, or more loosely applied to nodes regarded as being close to the root. Note that extant taxa that lie on branches connecting directly to the root are not more closely related to the root than any other extant taxa. While there must always be two or more equally "basal" clades sprouting from the root of every cladogram, those clades may differ widely in taxonomic rank, species diversity, or both. If ''C'' is a basal clade within ''D'' that has the lowest rank of all basal clades within ''D'', ''C'' may be described as ''the'' basal taxon of that rank within ''D''. The concept of a ' key innovation' implies some degree of correlation between evolutionary innovation and diversification. However, such a correlation does not make a given case predicable, so ancestral characters should not be imputed to th ...
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Holonema Westolli
''Holonema'' is an extinct genus of relatively large, barrel-shaped arthrodire placoderms that were found in oceans throughout the world from the Mid to Late Devonian, when the last species perished in the Frasnian-Fammian extinction event. Most species of the genus are known from fragments of their armor, but the Gogo Reef species, ''H. westolli'', is known from whole, articulated specimens. Holonema is thought to have been a benthic-dwelling fish, based on its cambered body shape, which performs better hydrodynamically when closer to the sea bed.https://sciencepress.mnhn.fr/sites/default/files/articles/pdf/g1999n1a4.pdf According to these specimens, species of ''Holonema'' lived by grazing on stony, horn-shaped, stromatolite-like algae called oncholite, apparently by snipping off the points with a specialized snout. Species ''H. rugosum'' ''H. rugosum'' is the type species. It was originally described as a species of the antiarch genus, ''Pterichthys''. Fossils are fou ...
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Groenlandaspis Antarcticus
''Groenlandaspis'' is an extinct genus of arthrodire from the Late Devonian. Fossils of the different species are found in late Devonian strata in all continents except eastern Asia. The generic name commemorates the fact that the first specimens of the type species were found in Greenland. Description As with all other arthrodires, ''Groenlandaspis'' had a joint in the back of its head with its thoracic armor, allowing for its head to be thrown back, increasing its gape. However, as its head is somewhat compressed in comparison with many other arthrodires, and as the dorsal side came to a low, pyramid-like peak, it is believed that ''Groenlandaspis'' could not crane its head back very far. It was a relatively small fish, only in length on average, though one unusually large species, ''G. riniensis'', reached almost a metre in length Gess, Robert W.; Whitfield, Alan K. (August 22, 2020). "Estuarine fish and tetrapod evolution: insights from a Late Devonian (Famennian) Gondwanan ...
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