Biological Pattern Formation
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Biological Pattern Formation
The science of pattern formation deals with the visible, ( statistically) orderly outcomes of self-organization and the common principles behind similar patterns in nature. In developmental biology, pattern formation refers to the generation of complex organizations of cell fates in space and time. The role of genes in pattern formation is an aspect of morphogenesis, the creation of diverse anatomies from similar genes, now being explored in the science of evolutionary developmental biology or evo-devo. The mechanisms involved are well seen in the anterior-posterior patterning of embryos from the model organism ''Drosophila melanogaster'' (a fruit fly), one of the first organisms to have its morphogenesis studied, and in the eyespots of butterflies, whose development is a variant of the standard (fruit fly) mechanism. Patterns in nature Examples of pattern formation can be found in biology, physics, and science, and can readily be simulated with computer graphics, as describe ...
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Morphogen
A morphogen is a substance whose non-uniform distribution governs the pattern of tissue development in the process of morphogenesis or pattern formation, one of the core processes of developmental biology, establishing positions of the various specialized cell types within a tissue. More specifically, a morphogen is a signaling molecule that acts directly on cells to produce specific cellular responses depending on its local concentration. Typically, morphogens are produced by source cells and diffuse through surrounding tissues in an embryo during early development, such that concentration gradients are set up. These gradients drive the process of differentiation of unspecialised stem cells into different cell types, ultimately forming all the tissues and organs of the body. The control of morphogenesis is a central element in evolutionary developmental biology (evo-devo). History The term was coined by Alan Turing in the paper "The Chemical Basis of Morphogenesis", where he ...
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Fir Wave
A fir wave is a set of alternating bands of fir trees in sequential stages of development, observed in forests on exposed mountain slopes in several areas, including northeastern North America and Japan. Fir waves develop by wave-regeneration following wind disturbance, and is one of various types of patterned vegetation. Formation Fir waves form by the ecological process of wave-regeneration. When a tree falls, a gap in the canopy is formed. This exposes trees at the leeward edge of the gap to greater wind. These trees are thus more likely to die from damage and desiccation than windward trees. These leeward trees eventually die, gradually expanding the gap downwind. At the same time, young trees start to grow in the wind shadow in the windward portion of the gap, protected from the high winds by the surviving trees. The combination of dying trees at the leeward edge and regenerating trees at the windward edge results in the propagation of the fir waves in the direction of the ...
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Tiger Bush
Tiger bush, or brousse tigrée in the French language, is a patterned vegetation community and ground consisting of alternating bands of trees, shrubs, or grass separated by bare ground or low herb cover, that run roughly parallel to contour lines of equal elevation. The patterns occur on low slopes in arid and semi-arid regions, such as in Australia, Sahelian West Africa, and North America. Due to the natural water harvesting capacity, many species in tiger bush usually occur only under a higher rainfall regime. Formation The alternating pattern arises from the interplay of hydrological, ecological, and erosional phenomena. In the regions where tiger bush is present, plant growth is water-limited - the shortage of rainfall prevents vegetation from covering the entire landscape. Instead, trees and shrubs are able to establish by either tapping soil moisture reserves laterally or by sending roots to deeper, wetter soil depths. By a combination of plant litter, root macropo ...
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Patterned Vegetation
Patterned vegetation is a vegetation community that exhibits distinctive and repetitive patterns. Examples of patterned vegetation include fir waves, tiger bush, and string bog. The patterns typically arise from an interplay of phenomena that differentially encourage plant growth or mortality. A coherent pattern arises because there is a strong directional component to these phenomena, such as wind in the case of fir waves, or surface runoff in the case of tiger bush. The regular patterning of some types of vegetation is a striking feature of some landscapes. Patterns can include relatively evenly spaced patches, parallel bands or some intermediate between those two. These patterns in the vegetation can appear without any underlying pattern in soil types, and are thus said to “self-organize” rather than be determined by the environment. Several of the mechanisms underlying patterning of vegetation have been known and studied since at least the middle of the 20th century, howev ...
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Tiger Bush Niger Corona 1965-12-31
The tiger (''Panthera tigris'') is the largest living cat species and a member of the genus '' Panthera''. It is most recognisable for its dark vertical stripes on orange fur with a white underside. An apex predator, it primarily preys on ungulates, such as deer and wild boar. It is territorial and generally a solitary but social predator, requiring large contiguous areas of habitat to support its requirements for prey and rearing of its offspring. Tiger cubs stay with their mother for about two years and then become independent, leaving their mother's home range to establish their own. The tiger was first scientifically described in 1758. It once ranged widely from the Eastern Anatolia Region in the west to the Amur River basin in the east, and in the south from the foothills of the Himalayas to Bali in the Sunda Islands. Since the early 20th century, tiger populations have lost at least 93% of their historic range and have been extirpated from Western and Central Asia, ...
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Slime Mould
Slime mold or slime mould is an informal name given to several kinds of unrelated eukaryotic organisms with a life cycle that includes a free-living single-celled stage and the formation of spores. Spores are often produced in macroscopic multicellular or multinucleate fruiting bodies which may be formed through aggregation or fusion. Slime molds were formerly classified as fungi but are no longer considered part of that kingdom. Although not forming a single monophyletic clade, they are grouped within the paraphyletic group Protista. More than 900 species of slime mold occur globally. Their common name refers to part of some of these organisms' life cycles where they can appear as gelatinous "slime". This is mostly seen with the Myxogastria, which are the only macroscopic slime molds. Most slime molds are smaller than a few centimetres, but some species may reach sizes up to several square metres and masses up to 20 kilograms. They feed on microorganisms that live in ...
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Bacterial Patterns
The formation of patterns in the growth of bacterial colonies has extensively been studied experimentally. Resulting morphologies appear to depend on the growth conditions. They include well known morphologies such as dense branched morphology (DBM) or diffusion-limited aggregation (DLA), but much complex patterns and temporal behaviour can be found. A large number of studies on pattern formation in bacterial colonies have been performed in ''Bacillus subtilis'' and in ''Proteus mirabilis''. Mathematical modeling of colony growth can reproduce the observed morphologies and the effect of environmental changes. Employed models include: * Reaction–diffusion system * Cellular automata Colonies of Bacillus subtilis Colonies of Bacillus subtilis on a Petri dish can grow under controlled conditions. By varying agar concentration (which permits the control of the hardness of the medium), and the nutrient A nutrient is a substance used by an organism to survive, grow, and reprod ...
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Cerebral Cortex
The cerebral cortex, also known as the cerebral mantle, is the outer layer of neural tissue of the cerebrum of the brain in humans and other mammals. The cerebral cortex mostly consists of the six-layered neocortex, with just 10% consisting of allocortex. It is separated into two cortices, by the longitudinal fissure that divides the cerebrum into the left and right cerebral hemispheres. The two hemispheres are joined beneath the cortex by the corpus callosum. The cerebral cortex is the largest site of neural integration in the central nervous system. It plays a key role in attention, perception, awareness, thought, memory, language, and consciousness. The cerebral cortex is part of the brain responsible for cognition. In most mammals, apart from small mammals that have small brains, the cerebral cortex is folded, providing a greater surface area in the confined volume of the cranium. Apart from minimising brain and cranial volume, cortical folding is crucial for the brain ...
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Elastic Instability
Elastic instability is a form of instability occurring in elastic systems, such as buckling of beams and plates subject to large compressive loads. There are a lot of ways to study this kind of instability. One of them is to use the method of incremental deformations based on superposing a small perturbation on an equilibrium solution. Single degree of freedom-systems Consider as a simple example a rigid beam of length ''L'', hinged in one end and free in the other, and having an angular spring attached to the hinged end. The beam is loaded in the free end by a force ''F'' acting in the compressive axial direction of the beam, see the figure to the right. Moment equilibrium condition Assuming a clockwise angular deflection \theta, the clockwise moment exerted by the force becomes M_F = F L \sin\theta. The moment equilibrium equation is given by F L \sin \theta = k_\theta \theta where k_\theta is the spring constant of the angular spring (Nm/radian). Assuming \theta is ...
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Alan Turing
Alan Mathison Turing (; 23 June 1912 – 7 June 1954) was an English mathematician, computer scientist, logician, cryptanalyst, philosopher, and theoretical biologist. Turing was highly influential in the development of theoretical computer science, providing a formalisation of the concepts of algorithm and computation with the Turing machine, which can be considered a model of a general-purpose computer. He is widely considered to be the father of theoretical computer science and artificial intelligence. Born in Maida Vale, London, Turing was raised in southern England. He graduated at King's College, Cambridge, with a degree in mathematics. Whilst he was a fellow at Cambridge, he published a proof demonstrating that some purely mathematical yes–no questions can never be answered by computation and defined a Turing machine, and went on to prove that the halting problem for Turing machines is undecidable. In 1938, he obtained his PhD from the Department of Mathemati ...
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