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Brochocoleus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †'' Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †'' Brochocoleus rostratus'' Pon ...
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Brochocoleus Maculatus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †'' Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †'' Brochocoleus rostratus'' P ...
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Brochocoleus Rostratus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †'' Brochocoleus rostratus'' Po ...
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Brochocoleus Keenani
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Pon ...
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Brochocoleus Planus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †'' Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Pono ...
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Brochocoleus Punctatus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †''Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †'' Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Ponom ...
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Brochocoleus Cossiphus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †'' Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †''Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Ponoma ...
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Brochocoleus Maximus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †''Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †'' Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †''Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Ponomar ...
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Brochocoleus Alatus
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †'' Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †''Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †''Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †''Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Ponomare ...
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Brochocoleus Tobini
''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major systematic revision in 2020. * †''Brochocoleus alatus'' Ponomarenko, 1994 Ulan-Argalant Formation, Mongolia, Barremian * †''Brochocoleus cossiphus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus keenani'' Jarzembowski, Yan, Wang & Zhang, 2013 Weald Clay, United Kingdom, Barremian * †''Brochocoleus maculatus'' (Whalley 1985) Charmouth Mudstone Formation, United Kingdom, Sinemurian * †''Brochocoleus maximus'' Jarzembowski, Yan, Wang & Zhang, 2013 Purbeck Group, United Kingdom, Berriasian * †''Brochocoleus planus'' Ponomarenko, 1994 Dzun-Bain Formation, Mongolia, Aptian * †''Brochocoleus punctatus'' Hong, 1982 (type species) Chijinbao Formation, China, Aptian * †''Brochocoleus rostratus'' Ponomaren ...
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Ommatidae
The Ommatidae are a family of beetles in the suborder Archostemata. The Ommatidae are considered the extant beetle family that has most ancestral characteristics. There are only seven extant species, confined to Australia and South America. However, the geographical distribution was much wider during the Mesozoic spanning across Eurasia and Australia, suggesting that they were widespread on Pangea. So far, over 26 extinct genera containing over 170 species of these beetles have been described. Three extant genera have been assigned to this family: ''Omma,'' '' Tetraphalerus'' and '' Beutelius''. The family is considered to be a subfamily of Cupedidae by some authors, but have been found to be more closely related to Micromalthidae in molecular phylogenies. A close relationship with Micromalthidae is supported by several morphological characters, including those of the mandibles and male genitalia. Due to their rarity, their ecology is obscure, it is likely that their larvae feed ...
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Pareuryomma
''Pareuryomma'' is an extinct genus of Ommatine beetle. It is known from three species, ''P. angustum, P. cardiobasis'' and P''. tylodes'', all known from the Aptian aged Yixian Formation of China. The genus was first described in 2012, and formerly included the species ''Pareuryomma ancistrodonta'' from the Middle Jurassic Jiulongshan Formation, China, which was transferred to the genus '' Omma'' in 2020. The species ''P. angustum'' was formerly thought to be part of ''Brochocoleus ''Brochocoleus'' is an extinct genus of beetles in the family Ommatidae, known from the Early Jurassic to the Early Late Cretaceous. 9 species are currently recognised, with many species being reassigned to other genera by Kirejtshuk's major sys ...''. The genus is characterised by "slender body, long and subparallelsided or slightly narrowed head with moderately prominent eyes and more or less prominent temples, narrow pronotum with more or less explanate lateral carinae, elytra with the veins mo ...
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Weald Clay
Weald Clay or the Weald Clay Formation is a Lower Cretaceous sedimentary rock unit underlying areas of South East England, between the North and South Downs, in an area called the Weald Basin. It is the uppermost unit of the Wealden Group of rocks within the Weald Basin, and the upper portion of the unit is equivalent in age to the exposed portion of the Wessex Formation on the Isle of Wight. It predominantly consists of thinly bedded mudstone. The un-weathered form is blue/grey, and the yellow/orange is the weathered form, it is used in brickmaking. The formation was deposited in lagoonal, lacustrine and alluvial conditions that varied from freshwater to brackish. The clay alternates with other subordinate lithologies, notably hard red-weathering beds of ironstone, limestone (Sussex Marble) and sandstones, notably including the calcareous sandstone unit referred to as the Horsham Stone. It has a gradual, conformable contact with the underlying Tunbridge Wells Sand Formation, a ...
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