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Branchial Heart
Branchial hearts are accessory pumps that supplement the action of the systemic heart in a cephalopod's body. They are myogenic in nature. Branchial hearts are always in pairs located at the base of the gills. Each branchial heart consists of a single chamber.Cephalopoda Glossary
Tree of Life web project.
They pump blood through the gills via the afferent branchial veins. Since they only circulate , branchial hearts function under predominantly conditions. Branchial hearts also appear to be involved in

Chtenopteryx Sicula2
''Chtenopteryx'' is a genus of small, muscular, midwater squid in the monotypic family Chtenopterygidae. Four species are presently recognized in the genus, but more are believed to exist. These squid occupy tropical to subtropical The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Geographical z ... waters, probably at depths of between 500 and 1,000 m during the day and near-surface waters at night. The genus contains bioluminescent species. Species * '' Chtenopteryx canariensis'' Salcedo-Vargas & Guerrero-Kommritz, 2000 * '' Chtenopteryx chuni'' * Pfeffer, 1912 * '' Chtenopteryx sepioloides'' Rancurel, 1970 * '' Chtenopteryx sicula'' (Vérany, 1851), comb-finned squid or toothed-fin squid The species listed above with an asterisk (*) is questionable and needs further study to determine if it ...
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Systemic Heart
The heart is a muscular organ in most animals. This organ pumps blood through the blood vessels of the circulatory system. The pumped blood carries oxygen and nutrients to the body, while carrying metabolic waste such as carbon dioxide to the lungs. In humans, the heart is approximately the size of a closed fist and is located between the lungs, in the middle compartment of the chest. In humans, other mammals, and birds, the heart is divided into four chambers: upper left and right atria and lower left and right ventricles. Commonly the right atrium and ventricle are referred together as the right heart and their left counterparts as the left heart. Fish, in contrast, have two chambers, an atrium and a ventricle, while most reptiles have three chambers. In a healthy heart blood flows one way through the heart due to heart valves, which prevent backflow. The heart is enclosed in a protective sac, the pericardium, which also contains a small amount of fluid. The wall of t ...
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Cephalopod
A cephalopod is any member of the molluscan class Cephalopoda (Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral body symmetry, a prominent head, and a set of arms or tentacles (muscular hydrostats) modified from the primitive molluscan foot. Fishers sometimes call cephalopods "inkfish", referring to their common ability to squirt ink. The study of cephalopods is a branch of malacology known as teuthology. Cephalopods became dominant during the Ordovician period, represented by primitive nautiloids. The class now contains two, only distantly related, extant subclasses: Coleoidea, which includes octopuses, squid, and cuttlefish; and Nautiloidea, represented by ''Nautilus'' and ''Allonautilus''. In the Coleoidea, the molluscan shell has been internalized or is absent, whereas in the Nautiloidea, the external shell remains. About 800 living species of cephalopods have been ident ...
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Myogenic
The myogenic mechanism is how arteries and arterioles react to an increase or decrease of blood pressure to keep the blood flow constant within the blood vessel. Myogenic response refers to a contraction initiated by the myocyte itself instead of an outside occurrence or stimulus such as nerve innervation. Most often observed in (although not necessarily restricted to) smaller resistance arteries, this 'basal' myogenic tone may be useful in the regulation of organ blood flow and peripheral resistance, as it positions a vessel in a preconstricted state that allows other factors to induce additional constriction or dilation to increase or decrease blood flow. The smooth muscle of the blood vessels reacts to the stretching of the muscle by opening ion channels, which cause the muscle to depolarize, leading to muscle contraction. This significantly reduces the volume of blood able to pass through the lumen, which reduces blood flow through the blood vessel. Alternatively when the s ...
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Ctenidium (mollusc)
A ctenidium is a respiratory organ or gill which is found in many molluscs. This structure exists in bivalves, cephalopods, Polyplacophorans (chitons), and in aquatic gastropods such as freshwater snails and marine snails. Some aquatic gastropods possess one ctenidium known as monopectinate and others have a pair of ctenidia known as bipectinate. A ctenidium is shaped like a comb or a feather, with a central part from which many filaments or plate-like structures protrude, lined up in a row. It hangs into the mantle cavity and increases the area available for gas exchange.Respiratory system
The apple snail. Retrieved 2012-04-20. The word is Latinized but is derived from the ''ktenidion'' which means "litt ...
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Venous Blood
Venous blood is deoxygenated blood which travels from the peripheral blood vessels, through the venous system into the right atrium of the heart. Deoxygenated blood is then pumped by the right ventricle to the lungs via the pulmonary artery which is divided in two branches, left and right to the left and right lungs respectively. Blood is oxygenated in the lungs and returns to the left atrium through the pulmonary veins. Venous blood is typically colder than arterial blood, and has a lower oxygen content and pH. It also has lower concentrations of glucose and other nutrients, and has higher concentrations of urea and other waste products. The difference in the oxygen content of arterial blood and venous blood is known as the arteriovenous oxygen difference. Most medical laboratory tests are conducted on venous blood, with the exception of arterial blood gas tests. Venous blood is obtained for lab work by venipuncture (also called phlebotomy), or by finger prick In medicine, s ...
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Hemocyanin
Hemocyanins (also spelled haemocyanins and abbreviated Hc) are proteins that transport oxygen throughout the bodies of some invertebrate animals. These metalloproteins contain two copper atoms that reversibly bind a single oxygen molecule (O2). They are second only to hemoglobin in frequency of use as an oxygen transport molecule. Unlike the hemoglobin in red blood cells found in vertebrates, hemocyanins are not confined in blood cells but are instead suspended directly in the hemolymph. Oxygenation causes a color change between the colorless Cu(I) deoxygenated form and the blue Cu(II) oxygenated form. Species distribution Hemocyanin was first discovered in ''Octopus vulgaris'' by Leon Fredericq in 1878. The presence of copper in molluscs was detected even earlier by Bartolomeo Bizio in 1833. Hemocyanins are found in the Mollusca and Arthropoda including cephalopods and crustaceans and utilized by some land arthropods such as the tarantula ''Eurypelma californicum'', the em ...
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Urine
Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excretion, excreted from the body through the urethra. Cell (biology), Cellular metabolism generates many by-products that are rich in nitrogen and must be clearance (medicine), cleared from the Circulatory system, bloodstream, such as urea, uric acid, and creatinine. These by-products are expelled from the body during urination, which is the primary method for excreting water-soluble chemicals from the body. A urinalysis can detect nitrogenous wastes of the mammalian body. Urine plays an important role in the earth's nitrogen cycle. In balanced ecosystems, urine fertilizes the soil and thus helps plants to grow. Therefore, Reuse of excreta, urine can be used as a fertilizer. Some animals use it to territory (animal)#Scent marking, mark their territories. Historically, aged or fermented urine (kn ...
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Nautiloid
Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rel ...
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Nautilus
The nautilus (, ) is a pelagic marine mollusc of the cephalopod family Nautilidae. The nautilus is the sole extant family of the superfamily Nautilaceae and of its smaller but near equal suborder, Nautilina. It comprises six living species in two genera, the type of which is the genus ''Nautilus''. Though it more specifically refers to species ''Nautilus pompilius'', the name chambered nautilus is also used for any of the Nautilidae. All are protected under CITES Appendix II. Depending on species, adult shell diameter is between 4 and 10 inches. Nautilidae, both extant and extinct, are characterized by involute or more or less convolute shells that are generally smooth, with compressed or depressed whorl sections, straight to sinuous sutures, and a tubular, generally central siphuncle.Kümmel, B. 1964. Nautiloidae-Nautilida, in the Treatise on Invertebrate Paleontology, Geological Society of America and Univ of Kansas Press, Teichert and Moore eds. Having survived relatively u ...
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Cardiology
Cardiology () is a branch of medicine that deals with disorders of the heart and the cardiovascular system. The field includes medical diagnosis and treatment of congenital heart defects, coronary artery disease, heart failure, valvular heart disease and electrophysiology. Physicians who specialize in this field of medicine are called cardiologists, a specialty of internal medicine. Pediatric cardiologists are pediatricians who specialize in cardiology. Physicians who specialize in cardiac surgery are called cardiothoracic surgeons or cardiac surgeons, a specialty of general surgery. Specializations All cardiologists study the disorders of the heart, but the study of adult and child heart disorders each require different training pathways. Therefore, an adult cardiologist (often simply called "cardiologist") is inadequately trained to take care of children, and pediatric cardiologists are not trained to treat adult heart disease. Surgical aspects are not included in cardiology ...
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