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Bolboparia
''Bolboparia'' is an extinct genus of eodiscinid agnostid trilobites. It lived during the late Lower Cambrian in what today Canada and the United States. Taxonomy ''Bolboparia'' is most closely related to ''Acidiscus'' and slightly more distantly to '' Stigmadiscus''. Description Like all Agnostida, ''Bolboparia'' is diminutive and the headshield (or cephalon) and tailshield (or pygidium The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is compo ...) are of approximately the same size (or isopygous) and outline. Like all Weymouthiidae, ''Bolboparia'' lacks eyes and rupture lines (or sutures). The short, downsloping glabella and the bulging cheeks give it a very peculiar aspect. Distribution * ''Bolboparia superba'' and ''B. elongata'' are both known from the Lower Cambrian of the Un ...
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Acidiscus
''Acidiscus'' Rasetti, 1966, is a genus of Eodiscinid trilobite belonging to the family Weymouthiidae Kobayashi T. (1943), Order Agnostida Salter (1864). It lived during the Botomian stage = late Lower Cambrian Stage 4 (upper of two stages subdividing the unnamed Series 2); the upper Botomian boundary corresponds to base of the Middle Cambrian, Miaolingian Series and Wuliuan stage. Etymology * ''A. birdi'' is named for Dr. John M. Bird who collected the holotype. * ''A. hexacanthus'' is derived from the Greek hexa "six" and acanthos "spine", for having two pairs of border spines in addition to one pair of genal spines. * ''A. theristes'' is named after a Greek in the Iliad (an epic poem written by Homer around 800 B.C.E.) who accused Agamemnon of greed and Achilles of cowardice during the Trojan War. Taxonomy A cladogram showing the relationship between several species of the genera ''Acidiscus'', '' Bolboparia'' and '' Stigmadiscus'' ''Acidiscus'' is most closely related to ...
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Stigmadiscus Cladogram
''Stigmadiscus'' Rasetti, 1966, is a genus of Lower Cambrian Eodiscinid trilobite belonging to the family Weymouthiidae The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zon ... Kobayashi (1943) Order Agnostida (Salter 1864) It lived during the Botomian stage, = late Lower Cambrian Stage 4 (upper of two stages subdividing the un-named Series 2); the upper Botomian Stage boundary corresponds to base of both the Middle Cambrian Wuliuan stage and Miaolingian Series. ''Stigmadiscus'' resembles ''Acidiscus'' in having a relatively short glabella, distinct S1 and S2 lateral furrows, and a slender occipital spine. In fact, according to Cotton & Fortey (2005, fig. 3) COTTON, T. J. and FORTEY, R. A. (2005). "5. Comparative morphology and relationships of the Agnostida". In Koenemann, S.; Jenner ...
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Weymouthiidae
The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zones in the former paleocontinents of Laurentia, Avalonia, Gondwana. The Weymouthiidae are all blind and lack free cheeks. Taxonomy Ancestors The Weymouthiidae have developed from a stock within the Hebediscidae The Hebediscidae Kobayashi, 1944, are a family of trilobites belonging to the order Agnostida that lived during the Lower Cambrian (Atdabanian to Toyonian). They are small or very small, and have a thorax of two or three segments. The Hebediscida .... Descendants The Weymouthiidae are a paraphyletic family because the Agnostina suborder is nested within it, particularly in the clade that further consists of the genera ''Mallagnostus'', ''Chelediscus'', ''Tannudiscus'' and ''Jinghediscus''. The trend in the Weymouthiid ...
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Stigmadiscus
''Stigmadiscus'' Rasetti, 1966, is a genus of Lower Cambrian Eodiscinid trilobite belonging to the family Weymouthiidae The Weymouthiidae Kobayashi 1943 are an extinct family of eodiscinid agnostid trilobites. They lived during the late Lower Cambrian and earliest Middle Cambrian (Botomian to Delamaran) in the so-called ''Olenellus''- and ''Eokochaspis''-zon ... Kobayashi (1943) Order Agnostida (Salter 1864) It lived during the Botomian stage, = late Lower Cambrian Stage 4 (upper of two stages subdividing the un-named Series 2); the upper Botomian Stage boundary corresponds to base of both the Middle Cambrian Wuliuan stage and Miaolingian Series. ''Stigmadiscus'' resembles ''Acidiscus'' in having a relatively short glabella, distinct S1 and S2 lateral furrows, and a slender occipital spine. In fact, according to Cotton & Fortey (2005, fig. 3) COTTON, T. J. and FORTEY, R. A. (2005). "5. Comparative morphology and relationships of the Agnostida". In Koenemann, S.; Jenner ...
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Animal
Animals are multicellular, eukaryotic organisms in the Kingdom (biology), biological kingdom Animalia. With few exceptions, animals Heterotroph, consume organic material, Cellular respiration#Aerobic respiration, breathe oxygen, are Motility, able to move, can Sexual reproduction, reproduce sexually, and go through an ontogenetic stage in which their body consists of a hollow sphere of Cell (biology), cells, the blastula, during Embryogenesis, embryonic development. Over 1.5 million Extant taxon, living animal species have been Species description, described—of which around 1 million are Insecta, insects—but it has been estimated there are over 7 million animal species in total. Animals range in length from to . They have Ecology, complex interactions with each other and their environments, forming intricate food webs. The scientific study of animals is known as zoology. Most living animal species are in Bilateria, a clade whose members have a Symmetry in biology#Bilate ...
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Fossils Of Canada
A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved in amber, hair, petrified wood and DNA remnants. The totality of fossils is known as the ''fossil record''. Paleontology is the study of fossils: their age, method of formation, and evolutionary significance. Specimens are usually considered to be fossils if they are over 10,000 years old. The oldest fossils are around 3.48 billion years old to 4.1 billion years old. Early edition, published online before print. The observation in the 19th century that certain fossils were associated with certain rock strata led to the recognition of a geological timescale and the relative ages of different fossils. The development of radiometric dating techniques in the early 20th century allowed scientists to quantitatively measure the absolut ...
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Cambrian Trilobites
The Cambrian Period ( ; sometimes symbolized Ꞓ) was the first geological period of the Paleozoic Era, and of the Phanerozoic Eon. The Cambrian lasted 53.4 million years from the end of the preceding Ediacaran Period 538.8 million years ago (mya) to the beginning of the Ordovician Period mya. Its subdivisions, and its base, are somewhat in flux. The period was established as "Cambrian series" by Adam Sedgwick, who named it after Cambria, the Latin name for 'Cymru' (Wales), where Britain's Cambrian rocks are best exposed. Sedgwick identified the layer as part of his task, along with Roderick Murchison, to subdivide the large "Transition Series", although the two geologists disagreed for a while on the appropriate categorization. The Cambrian is unique in its unusually high proportion of sedimentary deposits, sites of exceptional preservation where "soft" parts of organisms are preserved as well as their more resistant shells. As a result, our understanding of the Cambrian bi ...
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Agnostida Genera
Agnostida is an order of arthropod which have classically been seen as a group of highly modified trilobites, though some recent research has doubted this placement. Regardless, they appear to be close relatives as part of the Artiopoda. They are present in the Lower Cambrian fossil record along with trilobites from the Redlichiida, Corynexochida, and Ptychopariida orders, and were highly diverse throughout the Cambrian. Agnostidan diversity severely declined during the Cambrian-Ordovician transition, and the last agnostidans went extinct in the Late Ordovician. Systematics The Agnostida are divided into two suborders — Agnostina and Eodiscina — which are then subdivided into a number of families. As a group, agnostids are isopygous, meaning their pygidium is similar in size and shape to their cephalon. Most agnostid species were eyeless. The systematic position of the order Agnostida within the class Trilobita remains uncertain, and there has been continuing ...
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Trilobite
Trilobites (; meaning "three lobes") are extinct marine arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period () and they flourished throughout the lower Paleozoic before slipping into a long decline, when, during the Devonian, all trilobite orders except the Proetida died out. The last extant trilobites finally disappeared in the mass extinction at the end of the Permian about 252 million years ago. Trilobites were among the most successful of all early animals, existing in oceans for almost 270 million years, with over 22,000 species having been described. By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record. The stu ...
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Pygidium
The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.Shultz, J.W. (1990). Evolutionary Morphology And Phylogeny of Arachnida. Cladistics 6: 1–38. Chelicerates In arachnids, the pygidium is formed by reduction of the last three opisthosomal segments to rings where there is no distinction between tergites and sternites. A pygidium is present in Palpigradi, Amblypygi, Thelyphonida, Schizomida, Ricinulei and in the extinct order Trigonotarbida. It is also present in early fossil representatives of horseshoe crabs. Trilobites In trilobites, the pygidium can range from extremely small (much smaller than the head, or cephalon) to larger than the cephalon. They can be smooth, as in order ...
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Pygidium
The pygidium (plural pygidia) is the posterior body part or shield of crustaceans and some other arthropods, such as insects and the extinct trilobites. In groups other than insects, it contains the anus and, in females, the ovipositor. It is composed of fused body segments, sometimes with a tail, and separated from thoracic segments by an articulation.Shultz, J.W. (1990). Evolutionary Morphology And Phylogeny of Arachnida. Cladistics 6: 1–38. Chelicerates In arachnids, the pygidium is formed by reduction of the last three opisthosomal segments to rings where there is no distinction between tergites and sternites. A pygidium is present in Palpigradi, Amblypygi, Thelyphonida, Schizomida, Ricinulei and in the extinct order Trigonotarbida. It is also present in early fossil representatives of horseshoe crabs. Trilobites In trilobites, the pygidium can range from extremely small (much smaller than the head, or cephalon) to larger than the cephalon. They can be smooth, as in order ...
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Cephalon (arthropod Head)
The cephalon is the head section of an arthropod. It is a tagma, i.e., a specialized grouping of arthropod segments. The word cephalon derives from the Greek κεφαλή (kephalē), meaning "head". Insects In insects, ''head'' is a preferred term. The insect head consists of five segments, including three (the labial, maxillary and mandibular) necessary for food uptake, which are altogether known as the gnathocephalon and house the suboesophageal ganglion of the brain, as well as the antennal segment, and an ocular segment, as well as a non segmented fused section of the head where the archicerebrum is housed known as the acron. See also arthropod head problem. Chelicerates and crustaceans In chelicerates and crustaceans, the cephalothorax is derived from the fusion of the cephalon and the thorax, and is usually covered by a single unsegmented carapace. In relation with the arthropod head problem, phylogeny studies show that members of the Malacostraca class of cru ...
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