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Bembicosoma
''Bembicosoma'' is a genus of synziphosurine, a paraphyletic group of fossil chelicerate arthropods. ''Bembicosoma'' was regarded as part of the clade Planaterga. Fossils of the single and type species, ''B. pomphicus'', have been discovered in deposits of the Silurian period in the Pentland Hills, Scotland (in the United Kingdom). ''Bembicosoma'' had been tentatively assigned as an eurypterid (sea scorpion) before its synziphosurine affinities revealed. The prosoma of ''Bembicosoma'' covered by a short, semicircular carapace without evidence of ophthalmic ridges. Within the 10-segmented opisthosoma The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to a ..., the first segment is nearly as wide as the carapace but reduced in length while the second segment is the largest. The dorsal surfac ...
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Chelicerate
The subphylum Chelicerata (from New Latin, , ) constitutes one of the major subdivisions of the phylum Arthropoda. It contains the sea spiders, horseshoe crabs, and arachnids (including harvestmen, scorpions, spiders, solifuges, ticks, and mites, among many others), as well as a number of extinct lineages, such as the eurypterids (sea scorpions) and chasmataspidids. The Chelicerata originated as marine animals in the Middle Cambrian period; the first confirmed chelicerate fossils, belonging to '' Sanctacaris'', date from 508 million years ago. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species. Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered ...
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Synziphosurina
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and ''Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expre ...
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Planaterga
Planaterga is a clade of prosomapod euchelicerates including several synziphosurid genera (mainly bunodids and pseudoniscids) and the group Dekatriata (which in turn includes arachnids, chasmataspidids and eurypterid Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million yea ...s). Planaterga is defined by the opisthosoma with tergites broadest at third or fourth and lacking enlarged axial nodes, carapace (prosomal dorsal shield) with reduced genal spines, as well as somite VII (first opisthosomal segment) with reduced appendages and microtergite. References Middle Ordovician first appearances {{Chelicerata-stub ...
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Eurypterid
Eurypterids, often informally called sea scorpions, are a group of extinct arthropods that form the Order (biology), order Eurypterida. The earliest known eurypterids date to the Darriwilian stage of the Ordovician period 467.3 Myr, million years ago. The group is likely to have appeared first either during the Early Ordovician or Late Cambrian period. With approximately 250 species, the Eurypterida is the most diverse Paleozoic Chelicerata, chelicerate order. Following their appearance during the Ordovician, eurypterids became major components of marine faunas during the Silurian, from which the majority of eurypterid species have been described. The Silurian genus ''Eurypterus'' accounts for more than 90% of all known eurypterid specimens. Though the group continued to diversify during the subsequent Devonian period, the eurypterids were heavily affected by the Late Devonian extinction event. They declined in numbers and diversity until becoming extinct during the Permian–Tri ...
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Synziphosurine
Synziphosurina is a paraphyletic group of chelicerate arthropods previously thought to be basal horseshoe crabs (Xiphosura). It was later identified as a grade composed of various basal euchelicerates, eventually excluded form the monophyletic Xiphosura ''sensu stricto'' and only regarded as horseshoe crabs under a broader sense ('Xiphosura' ''sensu lato''). Synziphosurines survived at least since early Ordovician to early Carboniferous in ages, with most species are known from the in-between Silurian strata. File:20200920 Synziphosurina dorsal segmentation.png, Dorsal morphology and variation of opisthosomal segmentation across synziphosurines. While ''Weinbergina'' and ''Willwerathia'' being exceptionally large, most synziphosurines are small arthropods with body length ranging only about a few centimeters long. The body of synziphosurine composed of a prosoma covered by a dome-like carapace (prosomal dorsal shield) and an opisthosoma with usually unfused 9-11 segments expre ...
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Genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In the hierarchy of biological classification, genus comes above species and below family (taxonomy), family. In binomial nomenclature, the genus name forms the first part of the binomial species name for each species within the genus. :E.g. ''Panthera leo'' (lion) and ''Panthera onca'' (jaguar) are two species within the genus ''Panthera''. ''Panthera'' is a genus within the family Felidae. The composition of a genus is determined by taxonomy (biology), taxonomists. The standards for genus classification are not strictly codified, so different authorities often produce different classifications for genera. There are some general practices used, however, including the idea that a newly defined genus should fulfill these three criteria to be descriptively useful: # monophyly – all descendants ...
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United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and Northern Ireland. The United Kingdom includes the island of Great Britain, the north-eastern part of the island of Ireland, and many smaller islands within the British Isles. Northern Ireland shares a land border with the Republic of Ireland; otherwise, the United Kingdom is surrounded by the Atlantic Ocean, the North Sea, the English Channel, the Celtic Sea and the Irish Sea. The total area of the United Kingdom is , with an estimated 2020 population of more than 67 million people. The United Kingdom has evolved from a series of annexations, unions and separations of constituent countries over several hundred years. The Treaty of Union between the Kingdom of England (which included Wales, annexed in 1542) and the Kingdom of Scotland in 170 ...
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Silurian Arthropods
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago (Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during the ...
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Silurian First Appearances
The Silurian ( ) is a geologic period and system spanning 24.6 million years from the end of the Ordovician Period, at million years ago ( Mya), to the beginning of the Devonian Period, Mya. The Silurian is the shortest period of the Paleozoic Era. As with other geologic periods, the rock beds that define the period's start and end are well identified, but the exact dates are uncertain by a few million years. The base of the Silurian is set at a series of major Ordovician–Silurian extinction events when up to 60% of marine genera were wiped out. One important event in this period was the initial establishment of terrestrial life in what is known as the Silurian-Devonian Terrestrial Revolution: vascular plants emerged from more primitive land plants, dikaryan fungi started expanding and diversifying along with glomeromycotan fungi, and three groups of arthropods (myriapods, arachnids and hexapods) became fully terrestrialized. A significant evolutionary milestone during th ...
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Opisthosoma
The opisthosoma is the posterior part of the body in some arthropods, behind the prosoma (cephalothorax). It is a distinctive feature of the subphylum Chelicerata (arachnids, horseshoe crabs and others). Although it is similar in most respects to an abdomen (and is often referred to as such), the opisthosoma is differentiated by its inclusion of the respiratory organs (book lungs or book gills) and the heart. Segments The number of segments and appendages on the opisthosoma vary. Scorpions have 13, but the first is only seen during its embryological development. Other arachnids have fewer; harvestmen, for instance, have only ten. In general, appendages are absent or reduced, although in horseshoe crabs they persist as large plate-like limbs, called opercula or branchiophores, bearing the book gills, and that function in locomotion and gas exchange. In most chelicerates the opisthosomal limbs are greatly reduced and persist only as specialized structures, such as the silk-producing ...
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Carapace
A carapace is a Dorsum (biology), dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the underside is called the plastron. Crustaceans In crustaceans, the carapace functions as a protective cover over the cephalothorax (i.e., the fused head and thorax, as distinct from the abdomen behind). Where it projects forward beyond the eyes, this projection is called a rostrum (anatomy), rostrum. The carapace is Calcification, calcified to varying degrees in different crustaceans. Zooplankton within the phylum Crustacea also have a carapace. These include Cladocera, ostracods, and Isopoda, isopods, but isopods only have a developed "cephalic shield" carapace covering the head. Arachnids In arachnids, the carapace is formed by the fusion of prosomal tergites into a single Plate (animal anatomy), plate which carries the e ...
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Prosoma
The cephalothorax, also called prosoma in some groups, is a tagma of various arthropods, comprising the head and the thorax fused together, as distinct from the abdomen behind. (The terms ''prosoma'' and ''opisthosoma'' are equivalent to ''cephalothorax'' and ''abdomen'' in some groups.) The word ''cephalothorax'' is derived from the Greek words for head (, ') and thorax (, '). This fusion of the head and thorax is seen in chelicerates and crustaceans; in other groups, such as the Hexapoda (including insects), the head remains free of the thorax. In horseshoe crabs and many crustaceans, a hard shell called the carapace covers the cephalothorax. Arachnid anatomy Fovea The fovea is the centre of the cephalothorax and is located behind the head (only in spiders).Dalton, Steve (2008). ''Spiders; The Ultimate Predators''. A & C Black, London. P.p. 19. . It is often important in identification. It can be transverse or procurved Smith, A. M. (1990c). Baboon spiders: Tarantulas of Afr ...
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