Azhdarcho
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Azhdarcho
''Azhdarcho'' is a genus of azhdarchid pterosaur from the late Cretaceous Period of the Bissekty Formation (middle Turonian stage, about 92 million years ago) of Uzbekistan, as well as the Zhirkindek Formation of Kazakhstan. It is known from fragmentary remains including the distinctive, elongated neck vertebrae that characterizes members of the family Azhdarchidae, a family that includes many giant pterosaurs such as ''Quetzalcoatlus''. The name ''Azhdarcho'' comes from the Persian word '' azhdar'' (), a dragon-like creature in Persian mythology. The type species is ''Azhdarcho lancicollis''. The specific epithet ''lancicollis'' is derived from the Latin words ''lancea'' (meaning "lance" or "spear") and ''collum'' ("neck"). History The fossil remains of ''Azhdarcho'' were recovered in the Kyzyl Kum desert (from the Taykarshinskaya unit of the Bissekty Formation) by Lev A. Nesov during expeditions to Central Asia in 1974–1981. The type specimen, given the catalog number Р...
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Neoazhdarchia
Azhdarchoidea (or azhdarchoids) is a group of pterosaurs within the suborder Pterodactyloidea, more specifically within the group Ornithocheiroidea. Pterosaurs belonging to this group lived throughout the Early and Late Cretaceous periods, with one tentative member, ''Tendaguripterus'', that lived in the Late Jurassic period. The largest azhdarchoids include members of the family Azhdarchidae, examples of these are ''Quetzalcoatlus'', ''Hatzegopteryx'', and ''Arambourgiania''. The Azhdarchoidea has been recovered as either closely related to the Ctenochasmatoidea, as the sister taxon of the Pteranodontoidea within the Ornithocheiroidea, or within the Tapejaroidea, which in turn was also within the Ornithocheiroidea. Classification Azhdarchoidea was given a phylogenetic definition by David Unwin in 2003. Unwin defined the group as the most recent common ancestor of ''Quetzalcoatlus'' and '' Tapejara'', and all its descendants.Unwin, D. M., (2003). "On the phylogeny and evolutionar ...
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Azhdarchid Pterosaur Neck Lengths
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Azhdarchidae
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Azhdarchid
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Azhdarchinae
Azhdarchidae (from the Persian word , , a dragon-like creature in Persian mythology) is a family of pterosaurs known primarily from the Late Cretaceous Period, though an isolated vertebra apparently from an azhdarchid is known from the Early Cretaceous as well (late Berriasian age, about 140 million years ago). Azhdarchids included some of the largest known flying animals of all time, but smaller cat-size members have also been found. Originally considered a sub-family of Pteranodontidae, Nesov (1984) named the Azhdarchinae to include the pterosaurs ''Azhdarcho'', ''Quetzalcoatlus'', and ''Titanopteryx'' (now known as ''Arambourgiania''). They were among the last known surviving members of the pterosaurs, and were a rather successful group with a worldwide distribution. By the time of the end-Cretaceous mass extinction, most pterosaur families except for the Azhdarchidae disappear from the fossil record, but recent studies indicate a wealth of pterosaurian fauna, including pteran ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Quetzalcoatlus
''Quetzalcoatlus'' is a genus of pterosaur known from the Late Cretaceous period of North America (Maastrichtian stage); its members were among the largest known flying animals of all time. ''Quetzalcoatlus'' is a member of the Azhdarchidae, a family of advanced toothless pterosaurs with unusually long, stiffened necks. Its name comes from the Aztec feathered serpent god Quetzalcoatl. The type species is ''Q. northropi'', named by Douglas Lawson in 1975; the genus also includes the smaller species ''Q. lawsoni'', which was known for many years as an unnamed species before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021. Discovery and species The first ''Quetzalcoatlus'' fossils were discovered in Texas, United States, from the Maastrichtian Javelina Formation at Big Bend National Park (dated to around 68 million years ago) in 1971 by Douglas A. Lawson, then a geology graduate student from the Jackson School of Geosciences at the University of Texas ...
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Bissekty Formation
The Bissekty Formation (sometimes referred to as Bissekt) is a geologic formation and Lagerstätte which crops out in the Kyzyl Kum desert of Uzbekistan, and dates to the Late Cretaceous Period. Laid down in the mid to late Turonian, it is dated to about 92 to 90 Ma (million years ago). Description The lithology of the sediment largely consists of cross bedded sandstones with interbeds of massive sandstone, well cemented intraformational conglomerate, siltstones and mudstones. Most of the fossils are found as clasts within the conglomerates. Fossil content The Bissekty Formation is characterised by a mix of marine, brackish, freshwater, and terrestrial animal fossils. This stands in contrast the strictly marine fossils found in the underlying Dzheirantui Formation, and indicates that the Bissekty was formed during the regression of a saltwater sea. The coastline expanded inland again in the upper portion of the Bissekty, represented by a proportional increase of fully aquat ...
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Pteranodontidae
The Pteranodontidae are a family of large pterosaurs of the Cretaceous Period of North America and Africa. The family was named in 1876 by Othniel Charles Marsh. Pteranodontids had a distinctive, elongated crest jutting from the rear of the head (most famously seen in ''Pteranodon'' itself). The spectacularly-crested ''Nyctosaurus'' is sometimes included in this family, though usually placed in its own family, the Nyctosauridae (Nicholson & Lydekker, 1889). Modern researchers differ in their use of the concept. S. Christopher Bennett and Alexander Kellner have concluded that ''Nyctosaurus'' was not a pteranodontid. In 1994 Bennett defined a clade Pteranodontidae, also including species of the Anhangueridae. However, this definition has not been accepted by other workers. Alexander Kellner, for example, named several additional species for specimens previously classified as ''Pteranodon'', and placed ''P. sternbergi'' in a distinct genus, ''Geosternbergia''. Kellner re-defined Ptera ...
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Arambourgiania
''Arambourgiania'' is an extinct genus of azhdarchid pterosaur from the Late Cretaceous period (Maastrichtian stage) of Jordan, and possibly the United States.Harrell, T. Lynn Jr.; Gibson, Michael A.; Langston, Wann Jr. (2016). "A cervical vertebra of ''Arambourgiania philadelphiae'' (Pterosauria, Azhdarchidae) from the Late Campanian micaceous facies of the Coon Creek Formation in McNairy County, Tennessee, USA" ''Bull. Alabama Mus. Nat. Hist.'' 33:94–103 ''Arambourgiania'' was among the largest members of its family, the Azhdarchidae, and it is also one of the largest flying animals ever known. The incomplete left ulna of the "Sidi Chennane azhdarchid" from Morocco may have also belonged to ''Arambourgiania''. History of discovery In the early 1940s, a railway worker during repairs on the Amman-Damascus railroad near Russeifa found a two foot long fossil bone. In 1943 this was acquired by the director of a nearby phosphate mine, Amin Kawar, who brought it to the attention ...
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Lance Formation
The Lance (Creek) Formation is a division of Late Cretaceous (dating to about 69 - 66 Ma) rocks in the western United States. Named after Lance Creek, Wyoming, the microvertebrate fossils and dinosaurs represent important components of the latest Mesozoic vertebrate faunas. The Lance Formation is Late Maastrichtian in age (Lancian land mammal age), and shares much fauna with the Hell Creek Formation of Montana and North Dakota, the Frenchman Formation of southwest Saskatchewan, and the lower part of the Scollard Formation of Alberta. The Lance Formation occurs above the ''Baculites clinolobatus'' ammonite marine zone in Wyoming, the top of which has been dated to about 69 million years ago, and extends to the K-Pg boundary, 66 million years ago. However, the characteristic land vertebrate fauna of the Lancian age (which take its name from this formation) is only found in the upper strata of the Lance, roughly corresponding to the thinner equivalent formations such as the Hell C ...
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Zhirkindek Formation
The Zhirkindek Formation is a Late Cretaceous geologic formation in Kazakhstan. The primary lithology is sand with interbeds of clay and silt, and represents a coastal to coastal marine environment. The formation has produced numerous fossils, including Lindholmemydid and Trionychid cryptodires, indeterminate remains of dinosaurs (Ankylosauridae indet., Hadrosauroidea indet., Neoceratopsia indet., Sauropoda indet., Tyrannosauroidea indet., Ornithomimidae indet., Therizinosauroidea indet and Dromaeosauridae indet) and an indeterminate species of pterosaur ''Azhdarcho''. See also * List of dinosaur-bearing rock formations This list of dinosaur-bearing rock formations is a list of geologic formations in which dinosaur fossils have been documented. Containing body fossils * List of stratigraphic units with dinosaur body fossils ** List of stratigraphic units with few ... * List of fossiliferous stratigraphic units in Kazakhstan References {{reflist Geologic formations of K ...
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