Attercopus
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Attercopus
''Attercopus'' is an extinct genus of arachnids, containing one species ''Attercopus fimbriunguis'', known from flattened cuticle fossils from the Panther Mountain Formation in Upstate New York. It is placed in the extinct order Uraraneida, spider-like animals able to produce silk, but which lacked true spinnerets and retained a segmented abdomen bearing a flagellum-like tail resembling that of a whip scorpion. They are thought to be close to the origins of spiders. Its name is taken from the English dialect word ''attercop'' ("spider"), which came from ' ("poison-head"), from ' ("poison"), itself drawn from the Proto-Germanic ''*aitra-'' ("poisonous ulcer") and ''kopp-'' ("head"). In '' The Hobbit'', J. R. R. Tolkien had Bilbo use ''attercop'' to insult attacking spiders, the insult possibly deriving from its meaning in Northern England dialect of "peevish, ill-natured person". Cop or cob had also come to mean spider, as in cobweb. An important Early Devonian (about ) fos ...
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Uraraneida
Uraraneida is an extinct order of Paleozoic arachnids related to modern Spider, spiders. Two genera of fossils have been definitively placed in this order: ''Attercopus'' from the Devonian of United States and ''Permarachne'' from the Permian of Russia. Like spiders, they are known to have produced silk, but lack the characteristic Spinneret, spinnerets of modern spiders, and retain elongate Telson, telsons. Characteristics The first fossil now placed in the order was found in Gilboa, New York. In 1987, it was initially tentatively placed in the extinct order Trigonotarbida and named ''Gelasinotarbus''? ''fimbriunguis''. Later, partly on the basis of a supposed spinneret (spider), spinneret, it was identified as a spider and named ''Attercopus fimbriunguis''. Further specimens of this species were found, and when examined in detail, along with those assigned to the genus ''Permarachne'', features inconsistent with their placement as spiders were revealed. Silk producing spigots ...
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Spider
Spiders (order (biology), order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude spider silk, silk. They are the largest order of arachnids and rank seventh in total species diversity among all Order (biology), orders of organisms. Spiders are found worldwide on every continent except Antarctica, and have become established in nearly every land habitat. , 53,034 spider species in 136 Family (biology), families have been recorded by Taxonomy (biology), taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomy, Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segmentation (biology), segments are fused into two Tagma (biology), tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindr ...
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Panther Mountain Formation
The Panther Mountain Formation is a Formation (stratigraphy), geologic formation in New York (state), New York. It preserves fossils dating back to the Devonian Period (geology), period. It is located in the counties of Albany County, New York, Albany, Madison County, New York, Madison, Oneida County, New York, Oneida, Otsego County, New York, Otsego, and Schoharie County, New York, Schoharie. It is well known for its fossil arthropods preserved as flattened cuticles, including ''Attercopus'' and ''Dracochela''. Paleobiota Arachnids Myriapods Insects Eye fragments of supposed archaeognathan affinities, and scales of possible archaeognathan or zygentoman affinities are known. Flora See also * Gilboa Fossil Forest, forest existed in Givetian of New York * List of fossiliferous stratigraphic units in New York References

Devonian geology of New York (state) Devonian System of North America Givetian Stage Devonian southern paleotemperate deposits Shale fo ...
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Araneae
Spiders ( order Araneae) are air-breathing arthropods that have eight limbs, chelicerae with fangs generally able to inject venom, and spinnerets that extrude silk. They are the largest order of arachnids and rank seventh in total species diversity among all orders of organisms. Spiders are found worldwide on every continent except Antarctica, and have become established in nearly every land habitat. , 53,034 spider species in 136 families have been recorded by taxonomists. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900. Anatomically, spiders (as with all arachnids) differ from other arthropods in that the usual body segments are fused into two tagmata, the cephalothorax or prosoma, and the opisthosoma, or abdomen, and joined by a small, cylindrical pedicel. However, as there is currently neither paleontological nor embryological evidence that spiders ever had a se ...
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Permarachne
''Permarachne'' is an extinct genus of arachnids containing the single species ''Permarachne novokshonovi'' from the Permian ( Kungurian) of Russia, found in the Koshelevka Formation near the town of Suksun in Perm Krai Perm Krai (, ; ) is a federal subjects of Russia, federal subject of Russia (a Krais of Russia, krai), located in Eastern Europe. Its administrative center is Perm, Russia, Perm. The population of the krai was 2,532,405 (2021 Russian census, 2021 .... It is closely related to modern spiders but unlike them, it has a long thin tail, similar to its relative '' Attercopus'', it is known from the mostly complete holotype PIN 4909/12. It is about 1 cm in size. It initially was thought to be a spider, but is now thought to form a clade with at least its close relative ''Attercopus'', forming the grouping Uraraneida. References Permian arachnids Monotypic prehistoric arachnid genera Permian arthropods of Europe Kungurian genera {{paleo-arachnid-s ...
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Trigonotarbida
The order Trigonotarbida is a group of extinct arachnids whose fossil record extends from the late Silurian to the early Permian ( Pridoli to Sakmarian).Dunlop, J. A., Penney, D. & Jekel, D. 2020A summary list of fossil spiders and their relatives In World Spider Catalog. Natural History Museum Bern, online at http://wsc.nmbe.ch , version 20.5 These animals are known from several localities in Europe and North America, as well as a single record from Argentina. Trigonotarbids can be envisaged as spider-like arachnids, but without silk-producing spinnerets. They ranged in size from a few millimetres to a few centimetres in body length and had segmented abdomens (opisthosoma), with the dorsal exoskeleton ( tergites) across the backs of the animals' abdomens, which were characteristically divided into three or five separate plates. Probably living as predators on other arthropods, some later trigonotarbid species were quite heavily armoured and protected themselves with spines and tub ...
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Devonian Arthropods Of North America
The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of life on land occurred during the Devonian, as free- sporing land plants ( pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants ( pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrest ...
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Devonian Arachnids
The Devonian ( ) is a geologic period and system of the Paleozoic era during the Phanerozoic eon, spanning 60.3 million years from the end of the preceding Silurian period at million years ago ( Ma), to the beginning of the succeeding Carboniferous period at Ma. It is the fourth period of both the Paleozoic and the Phanerozoic. It is named after Devon, South West England, where rocks from this period were first studied. The first significant evolutionary radiation of life on land occurred during the Devonian, as free- sporing land plants (pteridophytes) began to spread across dry land, forming extensive coal forests which covered the continents. By the middle of the Devonian, several groups of vascular plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants ( pteridospermatophytes) appeared. This rapid evolution and colonization process, which had begun during the Silurian, is known as the Silurian-Devonian Terrestrial Revolu ...
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Science (journal)
''Science'' is the peer review, peer-reviewed academic journal of the American Association for the Advancement of Science (AAAS) and one of the world's top academic journals. It was first published in 1880, is currently circulated weekly and has a subscriber base of around 130,000. Because institutional subscriptions and online access serve a larger audience, its estimated readership is over 400,000 people. ''Science'' is based in Washington, D.C., United States, with a second office in Cambridge, UK. Contents The major focus of the journal is publishing important original scientific research and research reviews, but ''Science'' also publishes science-related news, opinions on science policy and other matters of interest to scientists and others who are concerned with the wide implications of science and technology. Unlike most scientific journals, which focus on a specific field, ''Science'' and its rival ''Nature (journal), Nature'' cover the full range of List of academ ...
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Russia
Russia, or the Russian Federation, is a country spanning Eastern Europe and North Asia. It is the list of countries and dependencies by area, largest country in the world, and extends across Time in Russia, eleven time zones, sharing Borders of Russia, land borders with fourteen countries. Russia is the List of European countries by population, most populous country in Europe and the List of countries and dependencies by population, ninth-most populous country in the world. It is a Urbanization by sovereign state, highly urbanised country, with sixteen of its urban areas having more than 1 million inhabitants. Moscow, the List of metropolitan areas in Europe, most populous metropolitan area in Europe, is the capital and List of cities and towns in Russia by population, largest city of Russia, while Saint Petersburg is its second-largest city and Society and culture in Saint Petersburg, cultural centre. Human settlement on the territory of modern Russia dates back to the ...
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Permian
The Permian ( ) is a geologic period and System (stratigraphy), stratigraphic system which spans 47 million years, from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.902 Mya. It is the sixth and last period of the Paleozoic Era; the following Triassic Period belongs to the Mesozoic Era. The concept of the Permian was introduced in 1841 by geologist Sir Roderick Murchison, who named it after the Perm Governorate, region of Perm in Russia. The Permian witnessed the diversification of the two groups of amniotes, the synapsids and the Sauropsida, sauropsids (reptiles). The world at the time was dominated by the supercontinent Pangaea, which had formed due to the collision of Euramerica and Gondwana during the Carboniferous. Pangaea was surrounded by the superocean Panthalassa. The Carboniferous rainforest collapse left behind vast regions of desert within the continental interior. Amniotes, which could better cope with these ...
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Chelicera
The chelicerae () are the mouthparts of the subphylum Chelicerata, an arthropod group that includes arachnids, horseshoe crabs, and sea spiders. Commonly referred to as "jaws", chelicerae may be shaped as either articulated fangs, or as a type of pincers. Some chelicerae, such as those found on nearly all spiders, are hollow and contain (or are connected to) venom glands, used to inject venom into prey or a perceived threat. Both pseudoscorpions and harvestmen have additional structures on their chelicerae that are used for grooming (papillae in pseudoscorpions, cheliceral teeth in Opiliones). In '' Paratrechalea'', males and females have shown to have a chelicerae dimorphism, because the chelicerae is used as a mating signal for females. Types Chelicerae can be divided into three kinds: jackknife chelicerae, scissor chelicerae, and three-segmented chelate chelicerae. Jackknife chelicerae The jackknife chelicera is subchelate (with fixed finger much reduced or absent) an ...
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