Archaeopterodactyloidea
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Archaeopterodactyloidea
Archaeopterodactyloidea (meaning "ancient Pterodactyloidea") is an extinct clade of pterodactyloid pterosaurs that lived from the middle Late Jurassic to the latest Early Cretaceous periods (Kimmeridgian to Albian stages) of Africa, Asia, Europe and North America. It was named by Alexander Wilhelm Armin Kellner in 1996 as the group that contains ''Germanodactylus'', ''Pterodactylus'', the Ctenochasmatidae and the Gallodactylidae. In 2003, Kellner defined the clade as a node-based taxon consisting of the last common ancestor of ''Pterodactylus'', ''Ctenochasma'' and ''Gallodactylus'' and all its descendants. Although phylogenetic analyses that based on David Unwin's 2003 analysis do not recover monophyletic Archaeopterodactyloidea, phylogenetic analyses that based on Kellner's analyses, or the analyses of Brian Andres (2008, 2010, 2018) recover monophyletic Archaeopterodactyloidea at the base of the Pterodactyloidea. The Antarctic Campanian specimen MN 7801-V was referred to Archa ...
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Germanodactylidae
Germanodactylidae is a controversial group of pterosaurs within the suborder Pterodactyloidea. It was first named by Yang Zhongjian in 1964, and given a formal phylogenetic definition in 2014 by Brian Andres, James Clark, and Xu Xing. They defined it as the least inclusive clade containing ''Germanodactylus cristatus'' and ''Normannognathus wellnhoferi'', which they considered to be close relatives at the time. However, more recent studies by the same researchers have found that these pterosaurs may be only distantly related. Studies performed in the 2000s suggested this group it contained three genera: ''Germanodactylus'', '' Normannognathus'' and ''Tendaguripterus''. Various studies have since placed these pterosaurs within the larger clades Archaeopterodactyloidea, Eupterodactyloidea, or Dsungaripteroidea, though it has also been recovered within the Ctenochasmatoidea. In several 2010s studies, the supposed "germanodactylid" species were not necessarily found to form a natur ...
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Pterosaur
Pterosaurs (; from Greek ''pteron'' and ''sauros'', meaning "wing lizard") is an extinct clade of flying reptiles in the order, Pterosauria. They existed during most of the Mesozoic: from the Late Triassic to the end of the Cretaceous (228 to 66 million years ago). Pterosaurs are the earliest vertebrates known to have evolved powered flight. Their wings were formed by a membrane of skin, muscle, and other tissues stretching from the ankles to a dramatically lengthened fourth finger. There were two major types of pterosaurs. Basal pterosaurs (also called 'non-pterodactyloid pterosaurs' or 'rhamphorhynchoids') were smaller animals with fully toothed jaws and, typically, long tails. Their wide wing membranes probably included and connected the hind legs. On the ground, they would have had an awkward sprawling posture, but the anatomy of their joints and strong claws would have made them effective climbers, and some may have even lived in trees. Basal pterosaurs were insectiv ...
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Euctenochasmatia
Euctenochasmatia is an extinct group of pterodactyloid pterosaurs. It was named by David Unwin in 2003 as the group that contains the most recent common ancestor of ''Pterodactylus'' and ''Ctenochasma'', and all their descendants. Euctenochasmatians were specialized pterosaurs that had elongated necks as well as specialized teeth. A peculiar family within this group is the Ctenochasmatidae, which most of the members had very distinguishing teeth that were lined within their elongated snouts. A genus called ''Pterofiltrus'' only had 112 teeth, but these teeth cover about 55.8% of the total skull, and the skull itself measured about in length. Description Euctenochasmatians had very distinctive features in comparison to other pterosaurs, including the shape of their jaws, as well as their highly specialized teeth. These teeth are thought to have been used for filter-feeding, the genus ''Pterodaustro'' for example, had a long snout and its lower jaws curve strongly upwards, and th ...
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Pterodactyloid
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like ''Nyctosaurus'' and ''Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. "Pteroda ...
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Germanodactylus
''Germanodactylus'' ("German finger") is a genus of germanodactylid pterodactyloid pterosaur from Upper Jurassic-age rocks of Germany, including the Solnhofen Limestone. Its specimens were long thought to pertain to '' Pterodactylus''. The head crest of ''Germanodactylus'' is a distinctive feature. History ''G. cristatus'' is based on specimen BSP 1892.IV.1, from the Solnhofen limestone of Eichstätt, Germany. It was originally described by Plieninger in 1901 as a specimen of ''Pterodactylus kochi'', and was given its current specific name by Carl Wiman in 1925, meaning "crested" in Latin. Yang Zhongjian determined that it deserved its own genus in 1964. A second species called ''G. ramphastinus'' (in 1858 accidentally revised to ''rhamphastinus'' by Christian Erich Hermann von Meyer) was named as a distinct species long before ''G. cristatus'', described by Johann Andreas Wagner in 1851 as a species of the now deprecated genus '' Ornithocephalus''. The specific name refe ...
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Pterodactyloidea
Pterodactyloidea (derived from the Greek words ''πτερόν'' (''pterón'', for usual ''ptéryx'') "wing", and ''δάκτυλος'' (''dáktylos'') "finger" meaning "winged finger", "wing-finger" or "finger-wing") is one of the two traditional suborders of pterosaurs ("wing lizards"), and contains the most derived members of this group of flying reptiles. They appeared during the middle Jurassic Period, and differ from the basal (though paraphyletic) rhamphorhynchoids by their short tails and long wing metacarpals (hand bones). The most advanced forms also lack teeth, and by the late Cretaceous, all known pterodactyloids were toothless. Many species had well-developed crests on the skull, a form of display taken to extremes in giant-crested forms like ''Nyctosaurus'' and ''Tupandactylus''. Pterodactyloids were the last surviving pterosaurs when the order became extinct at the end of the Cretaceous Period, together with the non-avian dinosaurs and most marine reptiles. "Pteroda ...
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Pterodactylus
''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of ''Pterodactylus'' have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of ''Pterodactylus'' have tentatively been identified from elsewhere in Europe and in Africa. ''Pterodactylus'' was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, ''Pterodactylus'' had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. ''Pterodactylus'' was a small pterosaur compared to other famo ...
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Gallodactylus
''Cycnorhamphus'' (meaning "swan beak") is a genus of gallodactylid ctenochasmatoid pterosaur from the Late Jurassic period of France and Germany, about 152 million years ago. It is probably synonymous with the genus ''Gallodactylus''. History In 1855, a fossil in a plate of shale from the Kimmeridgian, found near Nusplingen in Württemberg, holotype GPIT "Orig. Quenstedt 1855, Taf. 1" or GPIT 80, was named '' Pterodactylus Suevicus'' by Friedrich August Quenstedt. The specific name refers to the tribal area of Suevia. Quenstedt had earlier mentioned the find in a letter to Professor Heinrich Georg Bronn, which was published in 1854. In it he used the name ''Pterodactylus Württembergicus''. In 1855 and 1858, Christian Erich Hermann von Meyer adopted this older species name but it would be forgotten afterwards. The publication in 1854 was not meant to be a nomenclatural act. According to Peter Wellnhofer, ''Pterodactylus württembergicus'' should be considered a ''nomen obli ...
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Pterodactylus Antiquus
''Pterodactylus'' (from Greek () meaning 'winged finger') is an extinct genus of pterosaurs. It is thought to contain only a single species, ''Pterodactylus antiquus'', which was the first pterosaur to be named and identified as a flying reptile. Fossil remains of ''Pterodactylus'' have primarily been found in the Solnhofen limestone of Bavaria, Germany, which dates from the Late Jurassic period (early Tithonian stage), about 150.8 to 148.5 million years ago. More fragmentary remains of ''Pterodactylus'' have tentatively been identified from elsewhere in Europe and in Africa. ''Pterodactylus'' was a generalist carnivore that probably fed on a variety of invertebrates and vertebrates. Like all pterosaurs, ''Pterodactylus'' had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges. ''Pterodactylus'' was a small pterosaur compared to other famou ...
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Gallodactylidae
Gallodactylidae is a group of pterosaurs within the suborder Pterodactyloidea. Gallodactylids differed from other related pterosaurs in several distinct features, including fewer than 50 teeth present only in the jaw tips, and rounded crests present on the rear portion of the skull and jaws but not near the ends of their snouts. At least some species possessed jaw flanges, possibly used to bissect hard-shelled prey. History Gallodactylidae was named to contain ''Gallodactylus'' (now usually considered a synonym of ''Cycnorhamphus'') and its closest relatives. Many subsequent studies, however, showed that ''Gallodactylus'' did not form a clade with any non-synonymous pterosaurs that were not themselves part of a different family, and so the name was often ignored. The name returned to common use with the discovery of ''Gladocephaloideus'', a Chinese pterosaur species that shared many similarities with ''Cycnorhamphus''. Among other features, the Gallodactylidae was distinguished by ...
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Ctenochasmatidae
Ctenochasmatidae is a group of pterosaurs within the suborder Pterodactyloidea. They are characterized by their distinctive teeth, which are thought to have been used for filter-feeding. Ctenochasmatids lived from the Late Jurassic to the Early Cretaceous periods. The earliest known ctenochasmatid remains date to the Late Jurassic Kimmeridgian age. Previously, a fossil jaw recovered from the Middle Jurassic Stonesfield Slate formation in the United Kingdom, was considered the oldest known. This specimen supposedly represented a member of the family Ctenochasmatidae,Buffetaut, E. and Jeffrey, P. (2012). "A ctenochasmatid pterosaur from the Stonesfield Slate (Bathonian, Middle Jurassic) of Oxfordshire, England." ''Geological Magazine'', (advance online publication) though further examination suggested it actually belonged to a teleosaurid stem-crocodilian instead of a pterosaur. Classification Below is cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "cha ...
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Africa
Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area and 20% of its land area.Sayre, April Pulley (1999), ''Africa'', Twenty-First Century Books. . With billion people as of , it accounts for about of the world's human population. Africa's population is the youngest amongst all the continents; the median age in 2012 was 19.7, when the worldwide median age was 30.4. Despite a wide range of natural resources, Africa is the least wealthy continent per capita and second-least wealthy by total wealth, behind Oceania. Scholars have attributed this to different factors including geography, climate, tribalism, colonialism, the Cold War, neocolonialism, lack of democracy, and corruption. Despite this low concentration of wealth, recent economic expansion and the large and young population make Afr ...
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