Archaeognatha
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Archaeognatha
The Archaeognatha are an order of apterygotes, known by various common names such as jumping bristletails. Among extant insect taxa they are some of the most evolutionarily primitive; they appeared in the Middle Devonian period at about the same time as the arachnids. Specimens that closely resemble extant species have been found as both body and trace fossils (the latter including body imprints and trackways) in strata from the remainder of the Paleozoic Era and more recent periods. For historical reasons an alternative name for the order is Microcoryphia. Until the late 20th century the suborders Zygentoma and Archaeognatha comprised the order Thysanura; both orders possess three-pronged tails comprising two lateral cerci and a medial epiproct or ''appendix dorsalis''. Of the three organs, the appendix dorsalis is considerably longer than the two cerci; in this the Archaeognatha differ from the Zygentoma, in which the three organs are subequal in length. In the late 20th cen ...
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Apterygota
The name Apterygota is sometimes applied to a subclass of small, agile insects, distinguished from other insects by their lack of wings in the present and in their evolutionary history; notable examples are the silverfish, the firebrat, and the jumping bristletails. Their first known occurrence in the fossil record is during the Devonian period, 417–354 million years ago. The group Apterygota is not a clade; it is paraphyletic, and not universally recognized. As defined, the group contains two separate clades of wingless insects: Archaeognatha comprises jumping bristletails, while Zygentoma comprises silverfish and firebrats. The Zygentoma are in the clade Dicondylia with winged insects, a clade that includes all other insects, while Archaeognatha is sister to this lineage.A. Blanke, M. Koch, B. Wipfler, F. Wilde, B. Misof (2014) Head morphology of ''Tricholepidion gertschi'' indicates monophyletic Zygentoma. Frontiers in Zoology 11:16 doi:10.1186/1742-9994-11-16 The nymphs ...
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Insect
Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pairs of jointed legs, compound eyes and one pair of antennae. Their blood is not totally contained in vessels; some circulates in an open cavity known as the haemocoel. Insects are the most diverse group of animals; they include more than a million described species and represent more than half of all known living organisms. The total number of extant species is estimated at between six and ten million; In: potentially over 90% of the animal life forms on Earth are insects. Insects may be found in nearly all environments, although only a small number of species reside in the oceans, which are dominated by another arthropod group, crustaceans, which recent research has indicated insects are nested within. Nearly all insects hatch from eggs. ...
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Machilidae
The Machilidae are a family of insects belonging to the order Archaeognatha (the bristletails). There are around 250 described species worldwide. These insects are wingless, elongated and more or less cylindrical with a distinctive humped thorax and covered with tiny, close-fitting scales. The colour is usually grey or brown, sometimes intricately patterned. There are three "tails" at the rear of the abdomen: two cerci and a long central epiproct. They have large compound eyes, often meeting at a central point. They resemble the silverfish and the firebrat, which are from a different order, Zygentoma. Machilids undergo virtually no metamorphosis during their life cycles, and both nymphs and adults are generally inconspicuous herbivores and scavengers. Many species are restricted to rocky shorelines, but some are found in well-vegetated habitats inland. They can move very fast and often escape by jumping considerable distances when disturbed. Like all Archaeognatha, machilid ...
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Meinertellidae
The Meinertellidae are a small family of basal insects belonging to the order Archaeognatha. They are sometimes known as rock bristletails. These insects can be distinguished from members of the other Archaeognatha family, Machilidae, by the lack of scales at the base of the legs and antennae, head, and palps; along with possession of small abdominal sternites protruding slightly between the coxal plates. They can also be distinguished by patches of reddish to violet-brown hypodermal pigment on the appendages. Meinertellidae currently contains more than 170 species over 19 genera, clustered into the following groups: ''Machiloides, Machilinus, Hypomachiloides, Machilontus,'' and ''Meinertellus''. Distribution These insects are principally found in the southern hemisphere, and can be found in rain forests, regular forests, and on coastal cliffs. Taxonomy * †'' Cretaceobrevibusantennis'' Chen and Su 2017 Burmese amber, Myanmar, Cenomanian * †'' Cretaceomachilis'' St ...
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Dicondylia
The Dicondylia are a taxonomic group (taxon) that includes all insects except the jumping bristletails (Archaeognatha). Dicondylia have a mandible attached with two hinges to the head capsule (dicondyl), in contrast to a hypothetical ancestral mandible with a single ball joint (monocondyl); the members of Archaeognatha do in fact have dicondylic mandibles, though they are not identical to the structure seen in "true" dicondylic insects.Blanke, A., Machida, R., Szucsich, N.U., Wilde, F. and Misof, B. (2015) Mandibles with two joints evolved much earlier in the history of insects: dicondyly is a synapomorphy of bristletails, silverfish and winged insects. ''Syst. Entomol.'' 40: 357-364. https://doi.org/10.1111/syen.12107 Dicondyle mandible and other features The taxon is distinguished by the possession of a modified mandible with an additional joint canal, which also changes the muscle attachments of the mouth tools and allows a modified mandible movement compared to other mandib ...
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Thysanura
Thysanura is the now deprecated name of what was, for over a century, recognised as an order in the class Insecta. The two constituent groups within the former order, the Archaeognatha (jumping bristletails) and the Zygentoma (silverfish and firebrats), share several characteristics, such as of having three long caudal filaments, the lateral ones being the cerci, while the one between (telson) is a medial cerciform appendage, specifically an epiproct. They are also both wingless, and have bodies covered with fine scales, rather like the scales of the practically unrelated Lepidoptera. In the late 20th century, it was recognized that the two suborders were not sister taxa, therefore Thysanura was paraphyletic, and the two suborders were each raised to the status of an independent monophyletic order, with Archaeognatha sister taxon to the Dicondylia, including the Zygentoma. Although the group Thysanura is no longer recognized, the name still appears in some published material. An ...
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Zygentoma
Zygentoma are an order in the class Insecta, and consist of about 550 known species. The Zygentoma include the so-called silverfish or fishmoths, and the firebrats. A conspicuous feature of the order are the three long caudal filaments. The two lateral filaments are cerci, and the medial one is an epiproct or ''appendix dorsalis''. In this they resemble the Archaeognatha, although the cerci of Zygentoma, unlike in the latter order, are nearly as long as the epiproct. Until the late twentieth century the Zygentoma were regarded as a suborder of the Thysanura, until it was recognized that the order Thysanura was paraphyletic, thus raising the two suborders to the status of independent monophyletic orders, with Archaeognatha as sister group to the Dicondylia, including the Zygentoma. Etymology The name "Zygentoma" is derived from the Greek ('), in context meaning "yoke" or "bridge"; and ('), "insects", literally "cut into" because of the segmented anatomy of typical insects. pa ...
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Monura
Monura is an extinct suborder of wingless insects in the order Archaeognatha. They resembled their modern relatives, the silverfish, and had a single lengthy filament projecting from the end of the abdomen. They also had a pair of leg-like cerci and some non-ambulatory abdominal appendages. The largest specimens reached or more, not counting the length of the filament. Taxonomy * Suborder †Monura Sharov 1957 stat. nov. Carpenter 1992 ** Family † Dasyleptidae Sharov 1957 *** Genus †''Tonganoxichnus ''Tonganoxichnus'' (‘Tonganoxie, Kansas trace’) is a Pennsylvanian to Permian trace fossil that has been found in North America. The ichnogenus originally included two ichnospecies found in close association. ''T. buildexensis'' is interpre ...'' Mángano et al. 1997 (ichnotaxa) *** Genus †'' Dasyleptus'' Brongniart 1885 References Entognatha Carboniferous insects Pennsylvanian insects Permian insects Paleozoic insects of North America Carboniferous animal ...
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Epiproct
This glossary of entomology describes terms used in the formal study of insect species by entomologists. A–C A synthetic chlorinated hydrocarbon insecticide, toxic to vertebrates. Though its phytotoxicity is low, solvents in some formulations may damage certain crops. cf. the related Dieldrin, Endrin, Isodrin * D–F A synthetic chlorinated hydrocarbon insecticide, toxic to vertebrates. cf. the related Aldrin, Endrin, Isodrin A synthetic chlorinated hydrocarbon insecticide, toxic to vertebrates. Though its phytotoxicity is low, solvents in some formulations may damage certain crops. cf. the related Dieldrin, Aldrin, Isodrin G–L ...
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Ocelli
A simple eye (sometimes called a pigment pit) refers to a form of eye or an optical arrangement composed of a single lens and without an elaborate retina such as occurs in most vertebrates. In this sense "simple eye" is distinct from a multi-lensed "compound eye", and is not necessarily at all simple in the usual sense of the word. The structure of an animal's eye is determined by the environment in which it lives, and the behavioural tasks it must fulfill to survive. Arthropods differ widely in the habitats in which they live, as well as their visual requirements for finding food or conspecifics, and avoiding predators. Consequently, an enormous variety of eye types are found in arthropods. They possess a wide variety of novel solutions to overcome visual problems or limitations. Use of the term ''simple eye'' is flexible, and must be interpreted in proper context; for example, the eyes of humans and of other large animals such as most cephalopods, are ''camera eyes'' and ...
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Insect Mouthparts
Insects have mouthparts that may vary greatly across insect species, as they are adapted to particular modes of feeding. The earliest insects had chewing mouthparts. Most specialisation of mouthparts are for piercing and sucking, and this mode of feeding has evolved a number of times idependently. For example, mosquitoes and aphids (which are true bugs) both pierce and suck, however female mosquitoes feed on animal blood whereas aphids feed on plant fluids. Evolution Like most external features of arthropods, the mouthparts of Hexapoda are highly derived. Insect mouthparts show a multitude of different functional mechanisms across the wide diversity of insect species. It is common for significant homology to be conserved, with matching structures forming from matching primordia, and having the same evolutionary origin. However, even if structures are almost physically and functionally identical, they may not be homologous; their analogous functions and appearance might be the pr ...
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Compound Eye
A compound eye is a visual organ found in arthropods such as insects and crustaceans. It may consist of thousands of ommatidia, which are tiny independent photoreception units that consist of a cornea, lens, and photoreceptor cells which distinguish brightness and color. The image perceived by this arthropod eye is a combination of inputs from the numerous ommatidia, which are oriented to point in slightly different directions. Compared with single-aperture eyes, compound eyes have poor image resolution; however, they possess a very large view angle and the ability to detect fast movement and, in some cases, the polarization of light. Because a compound eye is made up of a collection of ommatidia, each with its own lens, light will enter each ommatidium instead of using a single entrance point. The individual light receptors behind each lens are then turned on and off due to a series of changes in the light intensity during movement or when an object in moving, creating a flic ...
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