Araripesaurus
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Araripesaurus
''Araripesaurus'' is a genus a pterosaur belonging to the suborder Pterodactyloidea, it was discovered in the Romualdo Formation of the Santana Group in northeastern Brazil, which dates back to the Aptian and Albian of the Early Cretaceous. The type species is ''A. castilhoi''. Discovery and naming The genus was named in 1971 by Brazilian paleontologist Llewellyn Ivor Price. The type species is ''Araripesaurus castilhoi''. The genus name refers to the Araripe Plateau. The specific name honors the collector Moacir Marques de Castilho, who in 1966 donated the chalk nodule containing the fossil. The holotype, DNPM (DGM 529-R), consists of a partial wing, including distal fragments of the radius and ulna, carpals, all metacarpals and several digits. The specimen was a subadult. Its wingspan was estimated at . Two other possible specimens are known; both consist of wing fragments and are roughly a third larger than the holotype, and were referred to the genus by Price. In 1985 Peter ...
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Ornithocheiridae
Ornithocheiridae (or ornithocheirids, meaning "bird hands") is a group of pterosaurs within the suborder Pterodactyloidea. These pterosaurs were among the last to possess teeth. Members that belong to this group lived from the Early to Late Cretaceous periods (Valanginian to Turonian stages), around 140 to 90 million years ago. Ornithocheirids are generally infamous for having an enormously controversial and very confusing taxonomy. Although agreements that these animals were related, and therefore similar to istiodactylids and pteranodontians, there is still no virtual consensus over the exact content and interrelationships of this group. Ornithocheirids were the most successful pterosaurs during their reign, they were also the largest pterosaurs before the appearance of the azhdarchids such as ''Quetzalcoatlus''. Ornithocheirids were excellent fish hunters, they used various flight techniques to catch their prey, and they are also capable of flying great distances without fl ...
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Anhanguera (pterosaur)
''Anhanguera'' () is a genus of pterodactyloid pterosaur known from the Early Cretaceous (Albian age, 125 to 112 million years ago) Romualdo Formation of Brazil and the Late Cretaceous ( Cenomanian age, 98 to 92.5 million years ago) Kem Kem Group of Morocco. This pterosaur is closely related to ''Ornithocheirus'', but belongs in the family Anhangueridae.Campos, D. de A., and Kellner, A. W. (1985). "Um novo exemplar de ''Anhanguera blittersdorffi'' (Reptilia, Pterosauria) da formação Santana, Cretaceo Inferior do Nordeste do Brasil." In Congresso Brasileiro de Paleontologia, Rio de Janeiro, Resumos, p. 13. The generic name comes from the Tupi words ''añanga'', meaning "spirit protector of the animals" + ''wera'' "bygone". Description ''Anhanguera'' was a fish-eating animal with a wingspan of about .Aureliano, T., Ghilardi, A. M., Duque, R. R., & Barreto, A. M. (2014). ON THE OCCURRENCE OF PTEROSAURIA IN EXU, PERNAMBUCO (LOWER CRETACEOUS ROMUALDO FORMATION, ARARIPE BASIN), N ...
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List Of Pterosaurs
This list of pterosaurs is a comprehensive listing of all genera that have ever been included in the order Pterosauria, excluding purely vernacular terms. The list includes all commonly accepted genera, but also genera that are now considered invalid, doubtful ('' nomen dubium''), or were not formally published ('' nomen nudum''), as well as junior synonyms of more established names, and genera that are no longer considered pterosaurian. The list currently includes 263 genera. Scope and terminology There is no official, canonical list of pterosaur genera, but the most thorough attempts can be found at the Pterosauria section of Mikko Haaramo's ''Phylogeny Archive'', the Genus Index at Mike Hanson's ''The Pterosauria'', supplemented by the Pterosaur Species List, and in the fourth supplement of Donald F. Glut's ''Dinosaurs: The Encyclopedia'' series. Authors and year The authors column lists the authors of the formal description responsible for the erection of the genus listed ...
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Timeline Of Pterosaur Research
This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs went extinct millions of years before humans evolved, humans have coexisted with pterosaur fossils for millennia. Before the development of paleontology as a formal science, these remains would have been interpreted through a mythological lens. Myths about thunderbirds told by the Native Americans of the modern Western United States may have been influenced by observations of ''Pteranodon'' fossils. These thunderbirds were said to have warred with water monsters, which agrees well with the co-occurrence of ''Pteranodon'' and the ancient marine reptiles of the seaway over which it flew. The formal study of pterosaurs began in the late 18th century when naturalist Cosimo Alessandro Collini of Mannheim, Germany published a description of ...
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Romualdo Formation
The Romualdo Formation is a geologic Konservat-Lagerstätte in northeastern Brazil's Araripe Basin where the states of Pernambuco, Piauí and Ceará come together. The geological formation, previously designated as the Romualdo Member of the Santana Formation, named after the village of Santana do Cariri, lies at the base of the Araripe Plateau. It was discovered by Johann Baptist von Spix in 1819. The strata were deposited during the Aptian stage of the Early Cretaceous in a lacustrine rift basin with shallow marine incursions of the proto-Atlantic. At that time, the South Atlantic was opening up in a long narrow shallow sea. The Romualdo Formation earns the designation of Lagerstätte due to an exceedingly well preserved and diverse fossil faunal assemblage. Some 25 species of fossil fishes are often found with stomach contents preserved, enabling paleontologists to study predator–prey relationships in this ecosystem. There are also fine examples of pterosaurs, reptiles a ...
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Santanadactylus
''Santanadactylus'' (meaning "Santana Formation finger") was a genus of pterodactyloid pterosaur from the Albian-age Romualdo Member of the Upper Cretaceous Santana Formation, of Barra do Jardim, Araripe Plateau, Ceará State, Brazil. Four species have been named, but today are no considered congeneric with each other. It was a rather large pterosaur. History and species The genus was named in 1980 by the Dutch paleontologist Paul de Buisonjé. The type species is ''S. brasilensis'', the specific name referring to Brazil. It is based on holotype UvA M 4894 ( Geological Institute of the University of Amsterdam), an upper part of the right humerus and a right scapulacoracoid. UvA M 4895, consisting of two cervical vertebrae from a different individual, was assigned as a paratype, referred because it was found in the same lot of 25 chalk nodules bought from collectors.De Buisonjé, P.H. (1980). ''Santanadactylus brasilensis'' nov. gen. nov. sp. a longnecked, large pterosaur from t ...
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Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ago), approximately. The Aptian succeeds the Barremian and precedes the Albian, all part of the Lower/Early Cretaceous. The Aptian partly overlaps the upper part of the Western European Urgonian Stage. The Selli Event, also known as OAE1a, was one of two oceanic anoxic events in the Cretaceous Period, which occurred around 120 Ma and lasted approximately 1 to 1.3 million years. The Aptian extinction was a minor extinction event hypothesized to have occurred around 116 to 117 Ma.Archangelsky, Sergio.The Ticó Flora (Patagonia) and the Aptian Extinction Event" ''Acta Paleobotanica'' 41(2), 2001, pp. 115-22. Stratigraphic definitions The Aptian was named after the small city of Apt in the Provence region of France, which is also known for its cry ...
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Metacarpals
In human anatomy, the metacarpal bones or metacarpus form the intermediate part of the skeleton, skeletal hand located between the phalanges of the fingers and the carpal bones of the wrist, which forms the connection to the forearm. The metacarpal bones are analogous to the metatarsal bones in the foot. Structure The metacarpals form a transverse arch to which the rigid row of distal carpal bones are fixed. The peripheral metacarpals (those of the thumb and little finger) form the sides of the cup of the palmar gutter and as they are brought together they deepen this concavity. The index metacarpal is the most firmly fixed, while the thumb metacarpal articulates with the trapezium and acts independently from the others. The middle metacarpals are tightly united to the carpus by intrinsic interlocking bone elements at their bases. The ring metacarpal is somewhat more mobile while the fifth metacarpal is semi-independent.Tubiana ''et al'' 1998, p 11 Each metacarpal bone consists o ...
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Morphology (biology)
Morphology is a branch of biology dealing with the study of the form and structure of organisms and their specific structural features. This includes aspects of the outward appearance (shape, structure, colour, pattern, size), i.e. external morphology (or eidonomy), as well as the form and structure of the internal parts like bones and organs, i.e. internal morphology (or anatomy). This is in contrast to physiology, which deals primarily with function. Morphology is a branch of life science dealing with the study of gross structure of an organism or taxon and its component parts. History The etymology of the word "morphology" is from the Ancient Greek (), meaning "form", and (), meaning "word, study, research". While the concept of form in biology, opposed to function, dates back to Aristotle (see Aristotle's biology), the field of morphology was developed by Johann Wolfgang von Goethe (1790) and independently by the German anatomist and physiologist Karl Friedrich Burdach ...
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Istiodactylus
''Istiodactylus'' is a genus of pterosaur that lived during the Early Cretaceous period, about 120 million years ago. The first fossil was discovered on the English Isle of Wight in 1887, and in 1901 became the holotype specimen of a new species, ''O. latidens'' (Latin for "wide tooth"), in the genus '' Ornithodesmus''. This species was moved to its own genus, ''Istiodactylus'', in 2001; this name is Greek for "sail finger". More specimens were described in 1913, and ''Istiodactylus'' was the only pterosaur known from three-dimensionally preserved fossils for much of the 20th century. In 2006, a species from China, ''I. sinensis'', was assigned to ''Istiodactylus'', but it has also been suggested to belong to a different genus. ''Istiodactylus'' was a large pterosaur; estimates of its wingspan range from . Its skull was about long, and was relatively short and broad for a pterosaur. The front of the snout was low and blunt, and bore a semicircle of 48 teeth. The triangular teeth ...
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Anhangueridae
Anhangueridae is a group of pterosaurs within the suborder Pterodactyloidea. They were among the last pterosaurs to possess teeth. A recent study discussing the group considered the Anhangueridae to be typified by a premaxillary crest and a lateral expansion in the distal rostrum. The same study presented a cladistic analysis, for which an "agreement subtree" was calculated. The Anhangueridae was found to be sister taxon to the large crested ''Tropeognathus''. Relationships There are competing theories of ornithocheiromorph phylogeny (evolutionary relationships). Below is cladogram following a topology recovered by Brian Andres, using the most recent iteration of his data set. The cladogram below follows Pêgas ''et al.'' (2019), who recovered Anhangueridae as a much more inclusive group. The analysis found most of the ornithocheirids falling into this family, while ''Ornithocheirus'' itself was recovered as a basal member of Ornithocheirae Ornithocheirae is an extinct clad ...
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Cladistics
Cladistics (; ) is an approach to biological classification in which organisms are categorized in groups (" clades") based on hypotheses of most recent common ancestry. The evidence for hypothesized relationships is typically shared derived characteristics ( synapomorphies'')'' that are not present in more distant groups and ancestors. However, from an empirical perspective, common ancestors are inferences based on a cladistic hypothesis of relationships of taxa whose character states can be observed. Theoretically, a last common ancestor and all its descendants constitute a (minimal) clade. Importantly, all descendants stay in their overarching ancestral clade. For example, if the terms ''worms'' or ''fishes'' were used within a ''strict'' cladistic framework, these terms would include humans. Many of these terms are normally used paraphyletically, outside of cladistics, e.g. as a 'grade', which are fruitless to precisely delineate, especially when including extinct species. R ...
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