Aneurophyton
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Aneurophyton
''Aneurophyton'' is a genus of extinct vascular plants that belong to the Aneurophytales, a class of progymnosperms. The genus is primarily known from records of two well-known species found in mostly Middle Devonian and Upper Devonian (late Eifelian to Famennian) outcrops in Belgium, China (West Junggar), Germany, and the United States (New York). Some uncertain species within the genus are also recorded from Middle Devonian outcrops in Kazakhstan, Russia (Timan and Siberia), and the Ukraine. While a number of species have been described in the paleobotanical literature, the genus likely only contains two well-circumscribed species, ''A. germanicum'' and ''A. doui'', and possibly a third species, ''A. olnense'', from Fammenian outcrops in Belgium. If the ages of the Early Devonian (Emsian) records of ''A. germanicum'' reported from Siberia are confirmed, these would constitute the oldest records of this genus. See also *Progymnosperm The progymnosperms are an extinct group o ...
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Progymnosperm
The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the seed plants, gymnosperms, ancestral to gymnosperms, acrogymnosperms and angiosperms (flowering plants). They have been treated formally at the rank (botany), rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as ''Protopteridium'', in which the vegetative organs consisted of relatively loose clusters of axes. ''Tetraxylopteris'' is another example of a genus lacking leaves. In more advanced aneurophytaleans such as ''Aneurophyton'' these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of pr ...
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Progymnosperm
The progymnosperms are an extinct group of woody, spore-bearing plants that is presumed to have evolved from the trimerophytes, and eventually gave rise to the seed plants, gymnosperms, ancestral to gymnosperms, acrogymnosperms and angiosperms (flowering plants). They have been treated formally at the rank (botany), rank of division Progymnospermophyta or class Progymnospermopsida (as opposite). The stratigraphically oldest known examples belong to the Middle Devonian order the Aneurophytales, with forms such as ''Protopteridium'', in which the vegetative organs consisted of relatively loose clusters of axes. ''Tetraxylopteris'' is another example of a genus lacking leaves. In more advanced aneurophytaleans such as ''Aneurophyton'' these vegetative organs started to look rather more like fronds, and eventually during Late Devonian times the aneurophytaleans are presumed to have given rise to the pteridosperm order, the Lyginopteridales. In Late Devonian times, another group of pr ...
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Vascular Plant
Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They also have a specialized non-lignified tissue (the phloem) to conduct products of photosynthesis. Vascular plants include the clubmosses, horsetails, ferns, gymnosperms (including conifers), and angiosperms (flowering plants). Scientific names for the group include Tracheophyta, Tracheobionta and Equisetopsida ''sensu lato''. Some early land plants (the rhyniophytes) had less developed vascular tissue; the term eutracheophyte has been used for all other vascular plants, including all living ones. Historically, vascular plants were known as "higher plants", as it was believed that they were further evolved than other plants due to being more complex organisms. However, this is an antiquated remnant of the obsolete scala naturae, and the term ...
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Middle Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Upper Devonian
The Devonian ( ) is a geologic period and system of the Paleozoic era, spanning 60.3 million years from the end of the Silurian, million years ago (Mya), to the beginning of the Carboniferous, Mya. It is named after Devon, England, where rocks from this period were first studied. The first significant adaptive radiation of life on dry land occurred during the Devonian. Free-sporing vascular plants began to spread across dry land, forming extensive forests which covered the continents. By the middle of the Devonian, several groups of plants had evolved leaves and true roots, and by the end of the period the first seed-bearing plants appeared. The arthropod groups of myriapods, arachnids and hexapods also became well-established early in this period, after starting their expansion to land at least from the Ordovician period. Fish reached substantial diversity during this time, leading the Devonian to often be dubbed the Age of Fishes. The placoderms began dominating al ...
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Eifelian
The Eifelian is the first of two faunal stages in the Middle Devonian Epoch. It lasted from 393.3 ± 1.2 million years ago to 387.7 ± 0.8 million years ago. It was preceded by the Emsian Stage and followed by the Givetian Stage. North American subdivisions of the Eifelian Stage include Southwood, and part of Cazenovia (or Cazenovian). Name and definition The Eifelian is named after the Eifel Mountains of Western Germany, which exposed the GSSP section at the Wetteldorf Richtschnitt outcrop. The base of the Eifelian is defined by the start of the '' Polygnathus partitus'' conodont zone. This layer lies within the Upper Heisdorf Formation, below the base of the Lauch Formation. Extinctions The end of the Eifelian was marked by a biological crisis known as the Kačák Event, a two-part interval of extinction which led to ecological turnover among ammonoids, conodonts, and other free-swimming animals. In deep marine waters, the event is indicated by anoxic black shales. Ther ...
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Famennian
The Famennian is the latter of two faunal stages in the Late Devonian Epoch. The most recent estimate for its duration estimates that it lasted from around 371.1 million years ago to 359.3 million years ago. An earlier 2012 estimate, still used by the International Commission on Stratigraphy, estimated that it lasted from million years ago to million years ago. It was preceded by the Frasnian stage and followed by the Tournaisian stage. Major events In the seas, a novel major group of ammonoid cephalopods called clymeniids appeared, underwent tremendous diversification and spread worldwide, then just as suddenly went extinct. The beginning of the Famennian is marked by the final stages of a major extinction event, the Kellwasser Event, which is the largest component of the Late Devonian Mass extinction. The end of the Famennian experiences a smaller but still quite severe extinction event, the Hangenberg Event. A brief episode of glaciation, possibly linked to the Hangenber ...
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Early Devonian
The Early Devonian is the first of three Epoch (geology), epochs comprising the Devonian period, corresponding to the Lower Devonian Series (stratigraphy), series. It lasted from and began with the Lochkovian Stage , which was followed by the Pragian from and then by the Emsian, which lasted until the Middle Devonian began, . During this time, the first Ammonoidea, ammonoids appeared, descending from Bactritida, bactritoid Nautiloidea, nautiloids. Ammonoids during this time period were simple and differed little from their nautiloid counterparts. These ammonoids belong to the order Agoniatitida, which in later epochs evolved to new ammonoid orders, for example Goniatite, Goniatitida and Clymeniida. This class of cephalopod molluscs would dominate the marine fauna until the beginning of the Mesozoic Era. References

{{Geological history Early Devonian, Geological epochs Devonian geochronology, *01 ...
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Emsian
The Emsian is one of three faunal stages in the Early Devonian Epoch. It lasted from 407.6 ± 2.6 million years ago to 393.3 ± 1.2 million years ago. It was preceded by the Pragian Stage and followed by the Eifelian Stage. It is named after the Ems river in Germany. The GSSP is located in the Zinzil'ban Gorge in the Kitab State Geological Reserve of Uzbekistan, above the contact with the Madmon Formation. In North America the Emsian Stage is represented by Sawkill or Sawkillian time. Biological events During this period, earliest known agoniatitid ammonoid fossils began appearing within this stage after first appearing in previous stage and began to evolutionarily radiate within this stage, in which a new ammonoid order Goniatitida rises in the end of Zlichovian stage (Siberian representation; corresponds to early Eifelian and after the end of Early Devonian, before 391.9 mya). Later agoniatitid ammonoids would die out in the Taghanic event in the upper middle Givetian ...
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Middle Devonian Plants
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