Amphistium
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Amphistium
''Amphistium paradoxum'' (from el, ἀμφί , 'on both sides', el, ιστίον 'sail', and el, παράδοξος 'extraordinary'), the only species classified under the genus ''Amphistium'', is a fossil fish which has been identified as a Paleogene relative of the flatfish, and as a transitional fossil. In a typical modern flatfish, the head is asymmetric with both eyes on one side of the head. In ''Amphistium'', the transition from the typical symmetric head of a vertebrate is incomplete, with one eye placed near the top of the head. ''Amphistium'' is among the many fossil fish species known from the Monte Bolca ''Lagerstätte'' of Lutetian Italy. ''Heteronectes ''Heteronectes chaneti'' is a fossil fish which has been identified as a primitive flatfish, dating to the early Eocene (Lutetian stage) of France. ''Heteronectes'' is reported to be a transitional fossil. In a typical modern flatfish, the head ...'' is a related, and very similar fossil from a slightly e ...
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Transitional Fossils
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Transitional Fossil
A transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation. Because of the incompleteness of the fossil record, there is usually no way to know exactly how close a transitional fossil is to the point of divergence. Therefore, it cannot be assumed that transitional fossils are direct ancestors of more recent groups, though they are frequently used as models for such ancestors. In 1859, when Charles Darwin's ''On the Origin of Species'' was first published, the fossil record was poorly known. Darwin described the perceived lack of transitional fossils as "the most obvious and gravest objection which can be ...
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Flatfish
A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating through or around the head during development. Some species face their left sides upward, some face their right sides upward, and others face either side upward. Many important food fish are in this order, including the flounders, soles, turbot, plaice, and halibut. Some flatfish can camouflage themselves on the ocean floor. Taxonomy Over 800 described species are placed into 16 families. Broadly, the flatfishes are divided into two suborders, Psettodoidei and Pleuronectoidei, with > 99% of the species diversity found within the Pleuronectoidei. The largest families are Soleidae, Bothidae and Cynoglossidae with more than 150 species each. There also exist two monotypic families ( Paralichthodidae and Oncopteridae). Some families ...
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Pleuronectiformes
A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating through or around the head during development. Some species face their left sides upward, some face their right sides upward, and others face either side upward. Many important food fish are in this order, including the flounders, soles, turbot, plaice, and halibut. Some flatfish can camouflage themselves on the ocean floor. Taxonomy Over 800 described species are placed into 16 families. Broadly, the flatfishes are divided into two suborders, Psettodoidei and Pleuronectoidei, with > 99% of the species diversity found within the Pleuronectoidei. The largest families are Soleidae, Bothidae and Cynoglossidae with more than 150 species each. There also exist two monotypic families (Paralichthodidae and Oncopteridae). Some families are ...
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Heteronectes
''Heteronectes chaneti'' is a fossil fish which has been identified as a primitive flatfish, dating to the early Eocene (Lutetian stage) of France. ''Heteronectes'' is reported to be a transitional fossil. In a typical modern flatfish, the head is asymmetric with both eyes on one side of the head. In ''Heteronectes'', the transition from the typical symmetric head of a vertebrate is incomplete, with one eye positioned near the top of the head, very similar (but less so) to its Italian relative '' Amphistium''. The rest of the skeleton also has some primitive features in common with other Percomorph groups, but absent in living flatfishes.Friedman M (2008). “The evolutionary origin of flatfish asymmetry”, ''Nature'' 454(7201): p. 209-212: The condition in modern, bottom-dwelling flatfish with both eyes on the same side of the head was cited by St. George Jackson Mivart as difficult to imagine how it could have evolved in a gradual fashion by natural selection, as proposed ...
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Paleogene
The Paleogene ( ; also spelled Palaeogene or Palæogene; informally Lower Tertiary or Early Tertiary) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period million years ago ( Mya) to the beginning of the Neogene Period Mya. It is the beginning of the Cenozoic Era of the present Phanerozoic Eon. The earlier term Tertiary Period was used to define the span of time now covered by the Paleogene Period and subsequent Neogene Period; despite no longer being recognised as a formal stratigraphic term, 'Tertiary' is still widely found in earth science literature and remains in informal use. Paleogene is often abbreviated "Pg" (but the United States Geological Survey uses the abbreviation PE for the Paleogene on the Survey's geologic maps). During the Paleogene, mammals diversified from relatively small, simple forms into a large group of diverse animals in the wake of the Cretaceous–Paleogene extinction event that ended the preceding ...
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National Geographic Society
The National Geographic Society (NGS), headquartered in Washington, D.C., United States, is one of the largest non-profit scientific and educational organizations in the world. Founded in 1888, its interests include geography, archaeology, and natural science, the promotion of environmental and historical conservation, and the study of world culture and history. The National Geographic Society's logo is a yellow portrait frame—rectangular in shape—which appears on the margins surrounding the front covers of its magazines and as its television channel logo. Through National Geographic Partners (a joint venture with The Walt Disney Company), the Society operates the magazine, TV channels, a website, worldwide events, and other media operations. Overview The National Geographic Society was founded on 13 January 1888 "to increase and diffuse geographic knowledge". It is governed by a board of trustees whose 33 members include distinguished educators, business exec ...
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Monte Bolca
Monte Bolca is a lagerstätte near Verona, Italy that was one of the first fossil sites with high quality preservation known to Europeans, and is still an important source of fossils from the Eocene. Geology Monte Bolca was uplifted from the Tethys Ocean floor during the formation of the Alps, in two stages: one 24 million years ago, and one between 30 and 50 million years ago. The entire formation consists of of limestone, all of which contain fossils, but interspersed in which are the lagerstätte layers that contain the highly preserved specimens. Within these layers, the fish and other specimens are so highly preserved that their organs are often completely intact in fossil form, and even the skin colorWilliams, MattTaphonomy of Monte Bolca University of Bristol can sometimes be determined. The normal rearrangement of the specimens caused by mud-dwelling organisms in the layer before it turned to stone has been avoided—it is assumed that the mud in question was low in o ...
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Lagerstätte
A Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that exhibits extraordinary fossils with exceptional preservation—sometimes including preserved soft tissues. These formations may have resulted from carcass burial in an anoxic environment with minimal bacteria, thus delaying the decomposition of both gross and fine biological features until long after a durable impression was created in the surrounding matrix. ''Lagerstätten'' span geological time from the Neoproterozoic era to the present. Worldwide, some of the best examples of near-perfect fossilization are the Cambrian Maotianshan shales and Burgess Shale, the Silurian Waukesha Biota, the Devonian Hunsrück Slates and Gogo Formation, the Carboniferous Mazon Creek, the Jurassic Posidonia Shale and Solnhofen Limestone, the Cretaceous Yixian, Santana, and Agua Nueva formations, the Eocene Green River Formation, the Miocene Foulden Maar ...
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Lutetian
The Lutetian is, in the geologic timescale, a stage or age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it is sometimes referred to as the Middle Eocene Subepoch. Stratigraphic definition The Lutetian was named after Lutetia, the Latin name for the city of Paris. The Lutetian Stage was introduced in scientific literature by French geologist Albert de Lapparent in 1883 and revised by A. Blondeau in 1981. The base of the Lutetian Stage is at the first appearance of the nanofossil ''Blackites inflatus'', according to an official reference profile ( GSSP) established in 2011. Of two candidates located in Spain, the Gorrondatxe section was chosen.See thwebsite of Eustoquio Molinafor these candidates. The top of the Lutetian (the base of the Bartonian) is at the first appearance of calcareous nanoplankton species ''Reticulofenestra reticulata''. The Lutetian overlaps with t ...
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Monotypic Ray-finned Fish Genera
In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispecific" or "monospecific" is sometimes preferred. In botanical nomenclature, a monotypic genus is a genus in the special case where a genus and a single species are simultaneously described. In contrast, an oligotypic taxon contains more than one but only a very few subordinate taxa. Examples Just as the term ''monotypic'' is used to describe a taxon including only one subdivision, the contained taxon can also be referred to as monotypic within the higher-level taxon, e.g. a genus monotypic within a family. Some examples of monotypic groups are: Plants * In the order Amborellales, there is only one family, Amborellaceae and there is only one genus, '' Amborella'', and in this genus there is only one species, namely ''Amborella trichopoda ...
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Prehistoric Ray-finned Fish Genera
Prehistory, also known as pre-literary history, is the period of human history between the use of the first stone tools by hominins 3.3 million years ago and the beginning of recorded history with the invention of writing systems. The use of symbols, marks, and images appears very early among humans, but the earliest known writing systems appeared 5000 years ago. It took thousands of years for writing systems to be widely adopted, with writing spreading to almost all cultures by the 19th century. The end of prehistory therefore came at very different times in different places, and the term is less often used in discussing societies where prehistory ended relatively recently. In the early Bronze Age, Sumer in Mesopotamia, the Indus Valley Civilisation, and ancient Egypt were the first civilizations to develop their own scripts and to keep historical records, with their neighbors following. Most other civilizations reached the end of prehistory during the following Iron Age. T ...
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