Amborella Trichopoda
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Amborella Trichopoda
''Amborella'' is a monotypic genus of understory shrubs or small trees endemic to the main island, Grande Terre, of New Caledonia in the southwest Pacific Ocean. The genus is the only member of the family Amborellaceae and the order Amborellales and contains a single species, ''Amborella trichopoda''. ''Amborella'' is of great interest to plant systematists because molecular phylogenetic analyses consistently place it as the sister group to all other flowering plants. Description ''Amborella'' is a sprawling shrub or small tree up to high. It bears alternate, simple evergreen leaves without stipules. The leaves are two-ranked, with distinctly serrated or rippled margins, and about long. ''Amborella'' has xylem tissue that differs from that of most other flowering plants. The xylem of ''Amborella'' contains only tracheids; vessel elements are absent. Xylem of this form has long been regarded as a primitive feature of flowering plants. The species is dioecious. This mea ...
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Aleksander Pavlovich Melikyan
Alexander is a male given name. The most prominent bearer of the name is Alexander the Great, the king of the Ancient Greek kingdom of Macedonia who created one of the largest empires in ancient history. Variants listed here are Aleksandar, Aleksander and Aleksandr. Related names and diminutives include Iskandar, Alec, Alek, Alex, Alexandre, Aleks, Aleksa and Sander; feminine forms include Alexandra, Alexandria, and Sasha. Etymology The name ''Alexander'' originates from the (; 'defending men' or 'protector of men'). It is a compound of the verb (; 'to ward off, avert, defend') and the noun (, genitive: , ; meaning 'man'). It is an example of the widespread motif of Greek names expressing "battle-prowess", in this case the ability to withstand or push back an enemy battle line. The earliest attested form of the name, is the Mycenaean Greek feminine anthroponym , , (/Alexandra/), written in the Linear B syllabic script. Alaksandu, alternatively called ''Alakasandu'' ...
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Sister Group
In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and taxon B are sister groups to each other. Taxa A and B, together with any other extant or extinct descendants of their most recent common ancestor (MRCA), form a monophyletic group, the clade AB. Clade AB and taxon C are also sister groups. Taxa A, B, and C, together with all other descendants of their MRCA form the clade ABC. The whole clade ABC is itself a subtree of a larger tree which offers yet more sister group relationships, both among the leaves and among larger, more deeply rooted clades. The tree structure shown connects through its root to the rest of the universal tree of life. In cladistic standards, taxa A, B, and C may represent specimens, species, genera, or any other taxonomic units. If A and B are at the same taxo ...
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Staminode
In botany, a staminode is an often rudimentary, sterile or abortive stamen, which means that it does not produce pollen.Jackson, Benjamin, Daydon; ''A Glossary of Botanic Terms with their Derivation and Accent''; Published by Gerald Duckworth & Co. London, 4th ed 1928 Staminodes are frequently inconspicuous and stamen-like, usually occurring at the inner whorl of the flower, but are also sometimes long enough to protrude from the corolla. Sometimes, the staminodes are modified to produce nectar, as in the Witch Hazel ''(Hamamelis)''.jin lu mei shu. Hamamelis Linnaeus, Sp. Pl. 1: 124. 1753. Flora of China 9: 32. 2003 Staminodes can be a critical characteristic for differentiating between species, for instance in the orchid genus ''Paphiopedilum'', and among the penstemons. In the case of Cannas, the petals are inconsequential and the staminodes are refined into eye-catching petal-like replacements. A spectacular example of staminode is given by ''Couroupita guianensis'', a ...
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Carpel
Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils'' and is typically surrounded by the pollen-producing reproductive organs, the stamens, collectively called the androecium. The gynoecium is often referred to as the "female" portion of the flower, although rather than directly producing female gametes (i.e. egg cells), the gynoecium produces megaspores, each of which develops into a female gametophyte which then produces egg cells. The term gynoecium is also used by botanists to refer to a cluster of archegonia and any associated modified leaves or stems present on a gametophyte shoot in mosses, liverworts, and hornworts. The corresponding terms for the male parts of those plants are clusters of antheridia within the androecium. Flowers that bear a gynoecium but no stamens are called ' ...
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Stamen
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the filament and an anther which contains '' microsporangia''. Most commonly anthers are two-lobed and are attached to the filament either at the base or in the middle area of the anther. The sterile tissue between the lobes is called the connective, an extension of the filament containing conducting strands. It can be seen as an extension on the dorsal side of the anther. A pollen grain develops from a microspore in the microsporangium and contains the male gametophyte. The stamens in a flower are collectively called the androecium. The androecium can consist of as few as one-half stamen (i.e. a single locule) as in '' Canna'' species or as many as 3,482 stamens which have been counted in the saguaro (''Carnegiea gigantea''). The androecium in ...
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Dioecious
Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is biparental reproduction. Dioecy has costs, since only about half the population directly produces offspring. It is one method for excluding self-fertilization and promoting allogamy (outcrossing), and thus tends to reduce the expression of recessive deleterious mutations present in a population. Plants have several other methods of preventing self-fertilization including, for example, dichogamy, herkogamy, and self-incompatibility. Dioecy is a dimorphic sexual system, alongside gynodioecy and androdioecy. In zoology In zoology, dioecious species may be opposed to hermaphroditic species, meaning that an individual is either male or female, in which case the synonym gonochory is more often used. Most animal species are dioeci ...
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Primitive (phylogenetics)
In phylogenetics, a primitive (or ancestral) character, trait, or feature of a lineage or taxon is one that is inherited from the common ancestor of a clade (or clade group) and has undergone little change since. Conversely, a trait that appears ''within'' the clade group (that is, is present in any subgroup within the clade but not all) is called advanced or derived. A clade is a group of organisms that consists of a common ancestor and all its lineal descendants. A primitive trait is the original condition of that trait in the common ancestor; advanced indicates a notable change from the original condition. These terms in biology contain no judgement about the sophistication, superiority, value or adaptiveness of the named trait. "Primitive" in biology means only that the character appeared first in the common ancestor of a clade group and has been passed on largely intact to more recent members of the clade. "Advanced" means the character has evolved within a later subgroup ...
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Vessel Element
A vessel element or vessel member (also called trachea or xylem vessel) is one of the cell types found in xylem, the water conducting tissue of plants. Vessel elements are typically found in angiosperms (flowering plants) but absent from most gymnosperms such as conifers. Vessel elements are the main feature distinguishing the " hardwood" of angiosperms from the " softwood" of conifers. Morphology Xylem is the tissue in vascular plants that conducts water (and substances dissolved in it) upwards in a plant. There are two kinds of cell that are involved in the actual transport: tracheids and vessel elements. Tracheids & vessel elements make the tracheary elements of xylem.Vessel elements are the building blocks of vessels, which constitute the major part of the water transporting system in those plants in which they occur. Vessels form an efficient system for transporting water (including necessary minerals) from the root to the leaves and other parts of the plant. In seco ...
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Tracheid
A tracheid is a long and tapered lignified cell in the xylem of vascular plants. It is a type of conductive cell called a tracheary element. Angiosperms use another type of tracheary element, called vessel elements, to transport water through the xylem. The main functions of tracheid cells are to transport water and inorganic salts, and to provide structural support for trees. There are often pits on the cell walls of tracheids, which allows for water flow between cells. Tracheids are dead at functional maturity and do not have a protoplast. The wood (softwood) of gymnosperms such as pines and other conifers is mainly composed of tracheids. Tracheids are also the main conductive cells in the primary xylem of ferns. The tracheid was first named after the German botanist Carl Gustav Sanio in 1863, from the German ''Tracheide''. Evolution Tracheids were the main conductive cells found in early vascular plants. In the first 140-150 million years of vascular plant evolution, tra ...
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Xylem
Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from the Ancient Greek word (''xylon''), meaning "wood"; the best-known xylem tissue is wood, though it is found throughout a plant. The term was introduced by Carl Nägeli in 1858. Structure The most distinctive xylem cells are the long tracheary elements that transport water. Tracheids and vessel elements are distinguished by their shape; vessel elements are shorter, and are connected together into long tubes that are called ''vessels''. Xylem also contains two other type of cells: parenchyma and fibers. Xylem can be found: * in vascular bundles, present in non-woody plants and non-woody parts of woody plants * in secondary xylem, laid down by a meristem called the vascular cambium in woody plants * as part of a stelar arrangement ...
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Stipule
In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many species they may be inconspicuous —or sometimes entirely absent, and the leaf is then termed ''exstipulate''. (In some older botanical writing, the term "stipule" was used more generally to refer to any small leaves or leaf-parts, notably prophylls.) The word ''stipule'' was coined by Linnaeus''Concise English Dictionary'' Wordsworth Editions Ltd. 1994, from Latin ''stipula'', straw, stalk. Types of stipules General characteristics The position of stipules on a plant varies widely from species to species, though they are often located near the base of a leaf. Stipules are most common on dicotyledons, where they appear in pairs alongside each leaf. Some monocotyledon plants display stipule-like structures, but only display one per lea ...
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Evergreen
In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, which completely lose their foliage during the winter or dry season. Evergreen species There are many different kinds of evergreen plants, both trees and shrubs. Evergreens include: *Most species of conifers (e.g., pine, hemlock, blue spruce, and red cedar), but not all (e.g., larch) *Live oak, holly, and "ancient" gymnosperms such as cycads *Most angiosperms from frost-free climates, and rainforest trees *All Eucalypts * Clubmosses and relatives * Bamboos The Latin binomial term , meaning "always green", refers to the evergreen nature of the plant, for instance :'' Cupressus sempervirens'' (a cypress) :''Lonicera sempervirens'' (a honeysuckle) :''Sequoia sempervirens'' (a sequoia) Leaf longevity in evergreen plants varies from a ...
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