Actinoceratida
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Actinoceratida
The Actinocerida are an order of generally straight, medium to large cephalopods that lived during the early and middle Paleozoic, distinguished by a siphuncle composed of expanded segments that extend into the adjacent chambers, in which deposits formed within contain a system of radial canals and a narrow space along the inner side of the connecting ring known as a paraspatium. (Teichert 1964) Septal necks are generally short and cyrtochoanitic, some being recumbent, some hook shaped. Most grew to lengths of about but some, like the Huroniidae of the Silurian grew significantly larger. Ecology The Actinocerida inhabited shallow to quite deep waters, where they alternated between swimming and lying on the bottom. They were predatory, and able to control their buoyancy to a greater degree than their contemporaries. Derivation The derivation of the Actinocerida remains enigmatic. They first appear late in the Early Ordovician (Cassinian Stage, late Floian) with the Georginida ...
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Actinoceras
''Actinoceras'' is the principal and root genus of the Actinoceratidae, a major family in the Actinocerida, that lived during the Middle and Late Ordovician. It is an extinct genus of nautiloid cephalopod that thrived in the warm waters of the United States during the Paleozoic era. Morphology ''Actinoceras'' are generally large, with typically straight shells reaching a meter or so in length (about 3 ft), with a blunt apex, and usually with a circular to subcircular cross section. . Shell characteristics The shells of ''Actinoceras'' are generally straight and long, although some are breviconic. Some are fusiform with the diameter decreasing from the anterior end of the phragmocone toward the aperture. Chambers are short and contain cameral deposits which are more concentrated apically and ventrally. Septa are close spaced, sutures are mostly transverse. The siphuncle, which varies in proportion to the size of the shell among species, is ventral, but not on the ventral ma ...
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Actinoceratidae
The Actinoceriatidae are a family of actinocerids named by Saemann in 1853 for those that grew to have large shells with blunt apices and large siphuncles with widely expanded segments and a generally arcuate endosiphucular canal system.Teichert 1964. Actinoceratoidea, Treatise on Invertebrate Paleontology, part K.(Nautiloidea) Their range is from the upper Middle Ordovician to the Lower Silurian. Actinocerids are generally straight-shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals Actinoceratids are derived from ''Wutinoceras'', possibly through an early '' Armenoceras'' or through '' Nybyoceras''Flower 1957.Studies of the Actinoceratida; Memoir 2; New Mexico Bureau of Mines and Mineral Resources, Socorro, NM and give rise to ''Lambeoceras'' and to the Huroniidae. Seven genera are included in the Actinoceratidae, ''Actinoceras'', '' Flowerocer ...
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Wutinoceratidae
The Wutinoceratidae are a family of early actinocerids defined by Shimazu and Obata in 1938Flower 1976. New American Wutinoceratidae with Review of Actinoceroid Occurrences in Eastern Hemisphere;PartI, Memoir 28; New Mexico Bureau of Mines and Mineral Resources for actinocerids with thick connecting rings and a complex irregular canal system. Actinocerids are generally straight shelled nautiloid cephalopods with a siphuncle composed of expanded segments, typically with thin connecting rings, in which the internal deposits are penetrated by a system of canals.Teichert 1964. Actinoceratoidea, Treatise on Invertebrate Paleontology, part K.(Nautiloidea) The Wutinoceratidae include three genera,Flower 1957. Studies of the Actinoceratida. Memoir 2; New Mexico Bureau of Mines and Mineral Resources, Socorro, NM ''Wutinoceras'', ''Cyrtonybyoceras'', and ''Adamsoceras'', known especially from the early Middle Ordovician (Whiterock stage) in northeastern China and North America, but foun ...
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Nautiloid
Nautiloids are a group of marine cephalopods ( Mollusca) which originated in the Late Cambrian and are represented today by the living ''Nautilus'' and ''Allonautilus''. Fossil nautiloids are diverse and speciose, with over 2,500 recorded species. They flourished during the early Paleozoic era, when they constituted the main predatory animals. Early in their evolution, nautiloids developed an extraordinary diversity of shell shapes, including coiled morphologies and giant straight-shelled forms ( orthocones). Only a handful of rare coiled species, the nautiluses, survive to the present day. In a broad sense, "nautiloid" refers to a major cephalopod subclass or collection of subclasses (Nautiloidea ''sensu lato''). Nautiloids are typically considered one of three main groups of cephalopods, along with the extinct ammonoids (ammonites) and living coleoids (such as squid, octopus, and kin). While ammonoids and coleoids are monophyletic clades with exclusive ancestor-descendant rel ...
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Lambeoceras
''Lambeoceras'' is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae. Morphological description ''Lambeoceras'' is of medium to moderately large size with a long, straight, depressed shell, broad in cross section with the dorsum and venter both about equally convex, meeting acutely along the sides. Chambers are short, and septa are closely spaced, forming broad lobes on the upper and lower sides, which meet in sharp saddles along the sides. The siphuncle is submarginal, near the ventral side and relatively narrow. Septal necks are extremely long, brims short and recumbent. Segments are broadly expanded, connecting rings thin. Radial canals within the siphuncle from broad arcs that may bifurcate close to the parispatium. Derivation and phylogeny ''Lambeoceras'' is derived from the same stock in '' Actinoceras'' that produced ''Kochoceras'' according to Flower ( ...
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Gonioceras
''Gonioceras'' is an extinct genus of actinocerid nautiloidean cephalopods typified by a broad, low shell; flattened ventrally, convexly rounded dorsally; top and bottom meeting at an acute angle along the sides. In most the shell is rather thin, especially along the lateral portion. The aperture is contracted. Sutures from broad ventral and dorsal lobes, more narrowly rounded ventro-lateral and dorso-lateral saddles, and sharp pointed lateral lobes; more complex than in later ''Lambeoceras''. The siphuncle is typically subcentral but may be closer to the venter; armenocerid in form with short segments and very short brims and containing a straight endosiphuncular canal system. ''Gonioceras'' is derived from ''Armenoceras'' and is the type genus of Gonioceratidae. Fossils are found in marine strata of the Chazy and Blackriveran of the Middle Ordovician in the Champlain region of eastern North America and at the Paquette Rapids of the Ottawa River in Ontario. The type species ...
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Adamsoceras
''Adamsoceras'' is a genus of actinocerids of the family Wutinoceratidae, with spheroidal siphuncle segments like '' Ormoceras'', but having a reticular canal system like ''Wutinoceras''. Adamsoceras has a slender, gently expanding, orthoconic shell that is slightly broader than high, i.e. depressed, with close spaced septa that form ventral lobes and a siphuncle that is near the ventral margin. ''Adamsoceras'' is known from rocks of Whiterockian age (early Middle Ordovician The Ordovician ( ) is a geologic period and system, the second of six periods of the Paleozoic Era. The Ordovician spans 41.6 million years from the end of the Cambrian Period million years ago (Mya) to the start of the Silurian Period Mya. T ...) in Nevada, the Baltic, Tasmania, and Manchuria. It may have been derived from ''Wutinoceras'', or from a common ancestor, and gave rise to ''Ormoceras''. The genotype is ''Adamsoceras isabelae'' from the upper Pogonip Group in Ikes Canyon in the Toquima Ran ...
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Lambeoceratidae
''Lambeoceras'' is a genus of large actinocerids with a convexly lenticular cross section from the Upper Ordovician of North America and the sole representative of the family Lambeoceratidae. Morphological description ''Lambeoceras'' is of medium to moderately large size with a long, straight, depressed shell, broad in cross section with the dorsum and venter both about equally convex, meeting acutely along the sides. Chambers are short, and septa are closely spaced, forming broad lobes on the upper and lower sides, which meet in sharp saddles along the sides. The siphuncle is submarginal, near the ventral side and relatively narrow. Septal necks are extremely long, brims short and recumbent. Segments are broadly expanded, connecting rings thin. Radial canals within the siphuncle from broad arcs that may bifurcate close to the parispatium. Derivation and phylogeny ''Lambeoceras'' is derived from the same stock in '' Actinoceras'' that produced ''Kochoceras'' according to Flower ( ...
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Bathmoceras
''Bathmoceras'' is a primitive cephalopod genus from the Middle and Upper Ordovician. It is a member of the order Cyrtocerinida and is the only genus in the family Bathmoceratidae. Derivation At one time ''Bathmoceras'' was thought to have possibly given rise to ''Polydesmia'', once thought to have been the ancestral actinocerid. Since then revision of actinocerid phylogeny and of Ordovician stratigraphy in east Asia have cast doubts on this hypothesis. ''Bathmoceras'' is most likely derived from ''Eothinoceras'' through lengthening of the septal necks and of the inward projections of the connecting rings in a forward direction. ''Eothinoceras'' differs in that the septal necks are vestigial rather than being somewhat long as in ''Bathmoceras'' and that the inward projections of the connecting rings point straight in rather than projecting forward. Both are straight shelled or slightly exogastric. Eothinoceras is also apparently more slender. Furnish and Glenister also include ...
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Canadian Epoch
The Canadian is the Lower or Early Ordovician in North America. The term is common in the older literature and has been well understood for more than a century. However it has no official recognition by the International Commission on Stratigraphy (ICS) and has been superseded by the more recently defined Ibexian series of western Utah. Background Dana introduced the Canadian as the name for a system separated from the rest of the Ordovician (Weller 1980), then known as the Lower Silurian, and referred to the rest of the Lower Silurian as the Trenton System. At that time the Ordovician had not yet been recognized. Later Ulrich redefined the Canadian as roughly equivalent to the Beekmantown strata of the Lower Ordovician. Flower (1957 p. 17) felt that recognition of the Canadian as a separate system would greatly solve problems in Early Paleozoic stratigraphy. As such, faunas in limestones of Canadian age are uniformly widespread and set off sharply from black shale graptolite ...
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Armenoceratidae
The Armenoceratidae are a family of early Paleozoic nautiloid cephalopods belonging to the order Actinocerida.Flower 1957.Studies of the Actinoceratida.; Memoir 2; New Mexico Bureau of Mines and Mineral Resources, Socorro, NM Teichert 1964. Actinoceratoidea, Treatise on Invertebrate Paleontology, part K.(Nautiloidea) The Armenoceratidae, established by Troedsson (1926) are characterized by large, straight, or slightly curved shells and large siphuncles with strongly expanded segments between the septa. Septal necks are short and abruptly recurved along brims. Radial canals in the endosiphuncular canal system are typically arched, curving forward and backward from near the septal foramina (openings) to connect with the parispatium on either side of the middle of each segments. The parispatium is the narrow opening between the inner side of the connecting rings in actinocerids and the internal siphuncular deposits that grow forward and back from the region of the septal open ...
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Armenoceras
''Armenoceras'' is a genus of actinocerid nautiloid cephalopods whose fossils ranged from the late Whiterockian Stage in the early Middle Ordovician, through the remainder of the period and on into the Upper Silurian. It is the type genus of the family Armenoceratidae. The shells of ''Armenoceras'' are straight and medium to large in size with a circular to subcircular cross section. The siphuncle is large, located subcentrally to resting on the ventral margin. Segments are wider than long, broadly expanded into the camerae. Septal necks short with wide brings that may be in contact with posterior surface of septa. In most the canal system within the siphuncle in is of the double arc type. Cameral deposits are rare. ''Armenoceras'' is derived from ''Wutinoceras'' through a thinning of the connecting rings and a simplification of the endosiphuncular canal system. The earliest ''Armenoceras'' known comes from the lower Whiterock equivalent in northern China and Korea but is unkn ...
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