Auxiliary Cell
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Auxiliary Cell
The auxiliary cell is a spore-like structure that form within the fungal family Gigasporaceae (order Gigasporales). Auxiliary cells have thin cell walls, (spiny), papillate, knobby or sometimes smooth surfaces, and are formed from hyphae after spore germination before the formation of mycorrhizae, and then on the extraradical hyphae in the soil. They may not be 'cells' in the biological sense of the word, as they are structures found with coenocytic hyphae belonging to members of the phylum (division) Glomeromycota. Mostly they are known from members of the Gigasporaceae. Currently this family contains '' Gigaspora'', '' Scutellospora'' and '' Racocetra'', but there are other generic names that have not been widely accepted (''Dentiscutata'', ''Cetraspora'', ''Fuscutata'' and ''Quatunica'') — all of these form auxiliary cells. Members of the genus '' Pacispora'' (another genus in the Diversisporales The ''Diversisporales'' are an order of generally hypogeous (underground ...
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Spore
In biology, a spore is a unit of sexual or asexual reproduction that may be adapted for dispersal and for survival, often for extended periods of time, in unfavourable conditions. Spores form part of the life cycles of many plants, algae, fungi and protozoa. Bacterial spores are not part of a sexual cycle, but are resistant structures used for survival under unfavourable conditions. Myxozoan spores release amoeboid infectious germs ("amoebulae") into their hosts for parasitic infection, but also reproduce within the hosts through the pairing of two nuclei within the plasmodium, which develops from the amoebula. In plants, spores are usually haploid and unicellular and are produced by meiosis in the sporangium of a diploid sporophyte. Under favourable conditions the spore can develop into a new organism using mitotic division, producing a multicellular gametophyte, which eventually goes on to produce gametes. Two gametes fuse to form a zygote which develops into a new s ...
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Fungus
A fungus ( : fungi or funguses) is any member of the group of eukaryotic organisms that includes microorganisms such as yeasts and molds, as well as the more familiar mushrooms. These organisms are classified as a kingdom, separately from the other eukaryotic kingdoms, which by one traditional classification include Plantae, Animalia, Protozoa, and Chromista. A characteristic that places fungi in a different kingdom from plants, bacteria, and some protists is chitin in their cell walls. Fungi, like animals, are heterotrophs; they acquire their food by absorbing dissolved molecules, typically by secreting digestive enzymes into their environment. Fungi do not photosynthesize. Growth is their means of mobility, except for spores (a few of which are flagellated), which may travel through the air or water. Fungi are the principal decomposers in ecological systems. These and other differences place fungi in a single group of related organisms, named the ''Eumycota'' (''true f ...
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Family (biology)
Family ( la, familia, plural ') is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between order and genus. A family may be divided into subfamilies, which are intermediate ranks between the ranks of family and genus. The official family names are Latin in origin; however, popular names are often used: for example, walnut trees and hickory trees belong to the family Juglandaceae, but that family is commonly referred to as the "walnut family". What belongs to a family—or if a described family should be recognized at all—are proposed and determined by practicing taxonomists. There are no hard rules for describing or recognizing a family, but in plants, they can be characterized on the basis of both vegetative and reproductive features of plant species. Taxonomists often take different positions about descriptions, and there may be no broad consensus across the scientific community for some time. The publishing of new data and opini ...
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Gigasporaceae
The Gigasporaceae are a family (biology), family of fungi in the order Diversisporales. Species in this family are widespread in distribution, and form arbuscular mycorrhiza in roots. A species under Gigasporaceae is ''Gigaspora gigantea''. The spores of ''G. gigantea'', found in specific sand dunes, commence in a healthy state of newly formed spores to dead and blackened in seven months through four identifiable steps: they begin as healthy greenish-yellow spores, turn into yellow with brown spots, then reddish-orange-brown, and ultimately dead. A cause of the symptoms of death in spores are soil organisms such as bacteria, protists, and microfauna. References

Diversisporales Fungus families {{Glomeromycota-stub ...
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Cell Wall
A cell wall is a structural layer surrounding some types of cells, just outside the cell membrane. It can be tough, flexible, and sometimes rigid. It provides the cell with both structural support and protection, and also acts as a filtering mechanism. Cell walls are absent in many eukaryotes, including animals, but are present in some other ones like fungi, algae and plants, and in most prokaryotes (except mollicute bacteria). A major function is to act as pressure vessels, preventing over-expansion of the cell when water enters. The composition of cell walls varies between taxonomic group and species and may depend on cell type and developmental stage. The primary cell wall of land plants is composed of the polysaccharides cellulose, hemicelluloses and pectin. Often, other polymers such as lignin, suberin or cutin are anchored to or embedded in plant cell walls. Algae possess cell walls made of glycoproteins and polysaccharides such as carrageenan and agar that are absent ...
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Hyphae
A hypha (; ) is a long, branching, filamentous structure of a fungus, oomycete, or actinobacterium. In most fungi, hyphae are the main mode of vegetative growth, and are collectively called a mycelium. Structure A hypha consists of one or more cells surrounded by a tubular cell wall. In most fungi, hyphae are divided into cells by internal cross-walls called "septa" (singular septum). Septa are usually perforated by pores large enough for ribosomes, mitochondria, and sometimes nuclei to flow between cells. The major structural polymer in fungal cell walls is typically chitin, in contrast to plants and oomycetes that have cellulosic cell walls. Some fungi have aseptate hyphae, meaning their hyphae are not partitioned by septa. Hyphae have an average diameter of 4–6 µm. Growth Hyphae grow at their tips. During tip growth, cell walls are extended by the external assembly and polymerization of cell wall components, and the internal production of new cell membrane. ...
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Mycorrhizae
  A mycorrhiza (from Greek μύκης ', "fungus", and ῥίζα ', "root"; pl. mycorrhizae, mycorrhiza or mycorrhizas) is a symbiotic association between a fungus and a plant. The term mycorrhiza refers to the role of the fungus in the plant's rhizosphere, its root system. Mycorrhizae play important roles in plant nutrition, soil biology, and soil chemistry. In a mycorrhizal association, the fungus colonizes the host plant's root tissues, either intracellularly as in arbuscular mycorrhizal fungi (AMF or AM), or extracellularly as in ectomycorrhizal fungi. The association is sometimes mutualistic. In particular species or in particular circumstances, mycorrhizae may have a parasitic association with host plants. Definition A mycorrhiza is a symbiotic association between a green plant and a fungus. The plant makes organic molecules such as sugars by photosynthesis and supplies them to the fungus, and the fungus supplies to the plant water and mineral nutrients, such as phosp ...
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Glomeromycota
Glomeromycota (often referred to as glomeromycetes, as they include only one class, Glomeromycetes) are one of eight currently recognized divisions within the kingdom Fungi, with approximately 230 described species. Members of the Glomeromycota form arbuscular mycorrhizas (AMs) with the thalli of bryophytes and the roots of vascular land plants. Not all species have been shown to form AMs, and one, ''Geosiphon pyriformis'', is known not to do so. Instead, it forms an endocytobiotic association with ''Nostoc'' cyanobacteria. The majority of evidence shows that the Glomeromycota are dependent on land plants (''Nostoc'' in the case of ''Geosiphon'') for carbon and energy, but there is recent circumstantial evidence that some species may be able to lead an independent existence. The arbuscular mycorrhizal species are terrestrial and widely distributed in soils worldwide where they form symbioses with the roots of the majority of plant species (>80%). They can also be found in wetla ...
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