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Australosyodon Nyaphuli
''Australosyodon'' is an extinct genus of dinocephalian therapsids from the middle Permian of South Africa. The first fossil was discovered in the 1980s near the village of Prince Albert Road in the Karoo region of South Africa. The genus is closely related to the genus ''Syodon'', found in the Russian Ischejewo fauna, and is thus regarded as the earliest example of an anteosaurid Dinocephalia outside of Russia. ''Australosyodon'' fossils have been recovered from the Eodicynodon Assemblage Zone, the lowest bank of the South African Beaufort Group, indicating the presence of primitive early therapsids in the southern hemisphere. The genus was first described by Bruce S. Rubidge in 1994. Description ''Australosyodon'' was a medium-sized dinocephalian. The skull was high and narrow, with a length of 26 cm, indicating a total body length of approximately 1.8 m (6 ft). The top of the skull displayed the thickening typical of dinocephalians. Overall there are many ...
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Middle Permian
The Guadalupian is the second and middle series/epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by the Lop ...
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Fang
A fang is a long, pointed tooth. In mammals, a fang is a modified maxillary tooth, used for biting and tearing flesh. In snakes, it is a specialized tooth that is associated with a venom gland (see snake venom). Spiders also have external fangs, which are part of the chelicerae. Fangs are most common in carnivores or omnivores, but some herbivores, such as fruit bats, have them as well. They are generally used to hold or swiftly kill prey, such as in large cats. Omnivorous animals, such as bears, use their fangs when hunting fish or other prey, but they are not needed for consuming fruit. Some apes also have fangs, which they use for threats and fighting. However, the relatively short canines of humans are not considered to be fangs. Fangs in religion, mythology and legend Certain mythological and legendary creatures such as dragons, gargoyles and yakshas are commonly depicted with prominent fangs. The fangs of vampires are one of their defining characteristics. The iconogra ...
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Anteosaurinae
Anteosaurinae is an extinct subfamily of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Syodontinae. Description These are very specialized, very large anteosaurs. The postcanine teeth are further reduced. Deepening of the postorbital region of the skull (behind the eyes) produced a larger temporal opening, indicating more muscle mass. The boss on the angular (rear of the jaw) has become very prominent, again, another sign of powerful jaw muscles. These huge animals were clearly formidable predators. In the Anteosaurinae, pachyostosis is taken to extremes. The dorsal (upper) surface of the nasal, frontal, and postfrontals (around and between/above the eyes) is thickened and rugose in the same manner as the tapinocephalids. Nevertheless these animals are too specialized and too late in time to have been the ancestors of the herbivorous tapinocephalids, so these characteristics evolved independently. Boonstra notes that the h ...
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Archaeosyodon Praeventor
''Archaeosyodon'' is an extinct genus of dinocephalian therapsids. It was medium-sized, reaching about 1.5–2 m (4–5 ft) in length. See also * List of therapsids This list of therapsids is an attempt to create a comprehensive listing of all genera that have ever been included in the Therapsida excluding mammals and purely vernacular terms. The list includes all commonly accepted genera, but also genera tha ... References Anteosaurs Prehistoric therapsid genera Permian synapsids of Asia Prehistoric synapsids of Europe Guadalupian synapsids Fossil taxa described in 1960 Guadalupian genus first appearances Guadalupian genus extinctions {{paleo-therapsid-stub ...
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Tapinocaninus Pamelae
''Tapinocaninus'' is an extinct genus of therapsids in the family Tapinocephalidae, of which it is the most basal member. Only one species is known, ''Tapinocaninus pamelae'' (meaning "Pam's humble canine"). Fossils have been found dating from the Middle Permian (Wordian age). Description This species is known from several skulls. It was a large animal, reaching up to in body mass. It is assumed to be either a herbivore or an omnivore. This dinocephalian has a heterodont dentition, consisting of incisors, canines and postcanines. Its canine teeth lack a definite crushing heel. The maxillary and premaxillary dentition is only partially preserved. The species is named in honour of Rubidge's mother. Discovery Fossils have been found in the Eodicynodon Assemblage Zone The ''Eodicynodon'' Assemblage Zone is a tetrapod assemblage zone or biozone which correlates to the Abrahamskraal Formation, Adelaide Subgroup of the Beaufort Group, a fossiliferous and geologically important geo ...
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Ulemosaurus Svijagensis
''Ulemosaurus'' is an extinct genus of dinocephalian therapsids that lived 265 to 260 million years ago, at Isheevo in Russian Tatarstan. It was a tapinocephalid, a group of bulky herbivores which flourished in the Middle Permian. ''Ulemosaurus'' and other tapinocephalians disappeared at the end of the Middle Permian. Description Only several partial skeletons and skulls have been found. The skull bones are extremely dense: about at its thickest. This thickening is possibly related to head-butting behavior, as some researchers suggest. The species is considered a herbivore, but because the mandible is heavily constructed some palaeontologists consider it a carnivore, with the species being able to use muscle power to cut prey up with its incisors. Classification ''Ulemosaurus'' is a large ''Moschops''-like form from Russia; it is probably similar enough to be included as a separate species of ''Moschops''. Despite its advanced characteristics, it lived slightly before the Karoo ...
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Estemmenosuchus Uralensis
''Estemmenosuchus'' (meaning "crowned crocodile" in Greek) is an extinct genus of large, early omnivorous therapsid. It is believed and interpreted to have lived during the middle part of the Middle Permian around 267 million years ago. The two species, ''E. uralensis'' and ''E. mirabilis'', are characterised by distinctive horn-like structures, which were probably used for intra-specific display. Both species of ''Estemmenosuchus'' are from the Perm (or Cis-Urals) region of Russia. Two other estemmenosuchids, ''Anoplosuchus'' and ''Zopherosuchus'', are now considered females of the species ''E. uralensis''. There were many complete and incomplete skeletons found together. Description ''Estemmenosuchus'' could reach a body length of more than . Its skull was long and massive, up to in length, and possessed several sets of large horns, somewhat similar to the antlers of a moose, growing upward and outward from the sides and top of the head. The animal had a sprawling posture as ...
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Dinocephalia
Dinocephalians (terrible heads) are a clade of large-bodied early therapsids that flourished in the Early and Middle Permian between 279.5 and 260 million years ago (Ma), but became extinct during the Capitanian mass extinction event. Dinocephalians included herbivorous, carnivorous, and omnivorous forms. Many species had thickened skulls with many knobs and bony projections. Dinocephalians were the first non-mammalian therapsids to be scientifically described and their fossils are known from Russia, China, Brazil, South Africa, Zimbabwe, and Tanzania. Description Apart from the Biarmosuchians, the dinocephalians are the least advanced therapsids, although still uniquely specialised in their own way. They retain a number of primitive characteristics (e.g. no secondary palate, small dentary) shared with their pelycosaur ancestors, although they are also more advanced in possessing therapsid adaptations like the expansion of the ilium and more erect limbs. They include carni ...
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Biarmosuchus Tener
''Biarmosuchus'' is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. ''Biarmosuchus'' was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was deposited by flood waters originating from the young Ural Mountains. Description ''Biarmosuchus'' was a medium-sized predator, similar in size to a large dog, grew up to 1.5–2 m in length with a skull length 15 cm (immature) to 21 cm. It was a lightly built, probably agile animal that would have fed on smaller tetrapods. Their legs are quite long, and the animals were probably quite agile in spite of their size. A large opening for the eye and a small temple opening common in primitive stem-mammals, this lends to a weak bite but how it ate is pure speculation. The teeth contained eight small incisors on the palate, followed by a canine tooth and a further five canine teeth. So together the species contained fourteen upper t ...
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Syodontinae
Syodontinae is a group of dinocephalian therapsids. It is one of two subfamilies in the family Anteosauridae, the other being Anteosaurinae. They are known from the Middle Permian Period of what is now Russia and South Africa. One of the best known syodontines is ''Syodon'' from Russia. The South African form ''Australosyodon'', is one of the earliest known Gondwanan anteosaurs. Description Syodontines lack the boss on the lower jaw that characterises the related anteosaurines, and they have often been considered more primitive in this respect. Classification Below is a cladogram showing syodontine relationships from a 2012 phylogenetic In biology, phylogenetics (; from Greek φυλή/ φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups o ... study of anteosaurians: References Anteosaurs Guadalupian first appearances Guada ...
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Titanophoneus
''Titanophoneus'' ("titanic murderer") is an extinct genus of carnivorous dinocephalian therapsid from the Middle Permian. It is classified within the family Anteosauridae. The type species is ''Titanophoneus potens''. Remains of ''Titanophoneus'' have been found at Isheevo in Russia. Description An adult skull would have reached 80 cm with a heavy long snout. The long tail and short limbs show the species to be a primitive therapsid unlike '' Inostrancevia'' which was more advanced. The structure of the limbs and the density of the bone are designed for a sprawling stance. The temporal opening is more advanced than Estemmenosuchidae but less advanced than ''Inostrancevia''. The teeth are large with 12 large palate incisors followed by 2 canines and various smaller back teeth. The lower palate is the same as the upper but without the canine teeth. The appearance of ''Titanophoneus'' is reminiscent to the sphenacodontid pelycosaurs, which included ''Dimetrodon''. Gallery im ...
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