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Archegonia
An archegonium (pl: archegonia), from the ancient Greek ''ἀρχή'' ("beginning") and ''γόνος'' ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female gamete. The corresponding male organ is called the antheridium. The archegonium has a long neck canal or venter and a swollen base. Archegonia are typically located on the surface of the plant thallus, although in the hornworts they are embedded. Bryophytes In bryophytes and other cryptogams sperm reach the archegonium by swimming in water films, whereas in Pinophyta and Angiosperms the pollen are delivered by wind or animal vectors and the sperm are delivered by means of a pollen tube. In the moss ''Physcomitrella patens'', archegonia are not embedded but are located on top of the leafy gametophore (s. Figure). The Polycomb protein FIE is expressed in the unfertilized egg cell (right) as the blue colour after GUS staining reveals. S ...
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Gametophore
Gametophores are prominent structures in seedless plants on which the reproductive organs are borne. The word gametophore and ‘-phore’ (Greek Φορά, "to be carried"). In mosses, liverworts and ferns (Archegoniata), the gametophores support gametangia (sex organs, female archegonia and male antheridia). If both archegonia and antheridia occur on the same plant, it is called monoecious. If there are separate female and male plants they are called dioecous. In Bryopsida the leafy moss plant (q. v. "Thallus") is the haploid gametophyte. It grows from its juvenile form, the protonema, under the influence of phytohormones (mainly cytokinins). Whereas the filamentous protonema grows by apical cell division, the gametophyte grows by division of three-faced apical cells. See also Rhizoid Rhizoids are protuberances that extend from the lower epidermal cells of bryophytes and algae. They are similar in structure and function to the root hairs of vascular land plants. Similar s ...
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Archegonium
An archegonium (pl: archegonia), from the ancient Greek ''ἀρχή'' ("beginning") and ''γόνος'' ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female gamete. The corresponding male organ is called the antheridium. The archegonium has a long neck canal or venter and a swollen base. Archegonia are typically located on the surface of the plant thallus, although in the hornworts they are embedded. Bryophytes In bryophytes and other cryptogams sperm reach the archegonium by swimming in water films, whereas in Pinophyta and Angiosperms the pollen are delivered by wind or animal vectors and the sperm are delivered by means of a pollen tube. In the moss ''Physcomitrella patens'', archegonia are not embedded but are located on top of the leafy gametophore (s. Figure). The Polycomb protein FIE is expressed in the unfertilized egg cell (right) as the blue colour after GUS staining reveals. S ...
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Archegonia
An archegonium (pl: archegonia), from the ancient Greek ''ἀρχή'' ("beginning") and ''γόνος'' ("offspring"), is a multicellular structure or organ of the gametophyte phase of certain plants, producing and containing the ovum or female gamete. The corresponding male organ is called the antheridium. The archegonium has a long neck canal or venter and a swollen base. Archegonia are typically located on the surface of the plant thallus, although in the hornworts they are embedded. Bryophytes In bryophytes and other cryptogams sperm reach the archegonium by swimming in water films, whereas in Pinophyta and Angiosperms the pollen are delivered by wind or animal vectors and the sperm are delivered by means of a pollen tube. In the moss ''Physcomitrella patens'', archegonia are not embedded but are located on top of the leafy gametophore (s. Figure). The Polycomb protein FIE is expressed in the unfertilized egg cell (right) as the blue colour after GUS staining reveals. S ...
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Moss
Mosses are small, non-vascular flowerless plants in the taxonomic division Bryophyta (, ) '' sensu stricto''. Bryophyta (''sensu lato'', Schimp. 1879) may also refer to the parent group bryophytes, which comprise liverworts, mosses, and hornworts. Mosses typically form dense green clumps or mats, often in damp or shady locations. The individual plants are usually composed of simple leaves that are generally only one cell thick, attached to a stem that may be branched or unbranched and has only a limited role in conducting water and nutrients. Although some species have conducting tissues, these are generally poorly developed and structurally different from similar tissue found in vascular plants. Mosses do not have seeds and after fertilisation develop sporophytes with unbranched stalks topped with single capsules containing spores. They are typically tall, though some species are much larger. ''Dawsonia'', the tallest moss in the world, can grow to in height. There are a ...
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Welwitschia
''Welwitschia'' is a monotypic gymnosperm genus, comprising solely the distinctive ''Welwitschia mirabilis'', endemic to the Namib desert within Namibia and Angola. ''Welwitschia'' is the only living genus of the family Welwitschiaceae and order Welwitschiales in the division Gnetophyta, and is one of three living genera in Gnetophyta, alongside ''Gnetum'' and ''Ephedra''. Informal sources commonly refer to the plant as a "living fossil". Naming ''Welwitschia'' is named after the Austrian botanist and doctor Friedrich Welwitsch, who described the plant in Angola in 1859. Welwitsch was so overwhelmed by the plant that he, "could do nothing but kneel down ..and gaze at it, half in fear lest a touch should prove it a figment of the imagination." Joseph Dalton Hooker of the Linnean Society of London, using Welwitsch's description and collected material along with material from the artist Thomas Baines who had independently recorded the plant in Namibia, described the species. Wel ...
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Gnetum
''Gnetum'' is a genus of gymnosperms, the sole genus in the family Gnetaceae within the Gnetophyta. They are tropical evergreen trees, shrubs and lianas. Unlike other gymnosperms, they possess vessel elements in the xylem. Some species have been proposed to have been the first plants to be insect-pollinated as their fossils occur in association with extinct pollinating scorpionflies.Ren D, Labandeira CC, Santiago-Blay JA, Rasnitsyn A, Shih CK, Bashkuev A, Logan MA, Hotton CL, Dilcher D. (2009). Probable Pollination Mode Before Angiosperms: Eurasian, Long-Proboscid Scorpionflies. Science, 326 (5954), 840-847. Molecular phylogenies based on nuclear and plastid sequences from most of the species indicate hybridization among some of the Southeast Asian species. Fossil-calibrated molecular-clocks suggest that the ''Gnetum'' lineages now found in Africa, South America and Southeast Asia are the result of ancient long-distance dispersal across seawater.Won H, Renner SS: The internal tran ...
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Angiosperm
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants that produce their seeds enclosed within a fruit. They are by far the most diverse group of land plants with 64 orders, 416 families, approximately 13,000 known genera and 300,000 known species. Angiosperms were formerly called Magnoliophyta (). Like gymnosperms, angiosperms are seed-producing plants. They are distinguished from gymnosperms by characteristics including flowers, endosperm within their seeds, and the production of fruits that contain the seeds. The ancestors of flowering plants diverged from the common ancestor of all living gymnosperms before the end of the Carboniferous, over 300 million years ago. The closest fossil relatives of flowering plants are uncertain and contentious. The earliest angiosperm fossils are in the ...
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Gymnosperm
The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, ''Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμνόσπερμος ( el, γυμνός, translit=gymnos, lit=naked, label=none and el, σπέρμα, translit=sperma, lit=seed, label=none), literally meaning 'naked seeds'. The name is based on the unenclosed condition of their seeds (called ovules in their unfertilized state). The non-encased condition of their seeds contrasts with the seeds and ovules of flowering plants (angiosperms), which are enclosed within an ovary. Gymnosperm seeds develop either on the surface of scales or leaves, which are often modified to form cones, or solitary as in yew, ''Torreya'', ''Ginkgo''. Gymnosperm lifecycles involve alternation of generations. They have a dominant diploid sporophyte phase and a reduced haploid gametophyte phase which is dependent on ...
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Ralf Reski
Ralf Reski (born 18 November 1958 in Gelsenkirchen) is a German professor of plant biotechnology and former dean of the Faculty of Biology of the University of Freiburg. He is also affiliated to the French École supérieure de biotechnologie Strasbourg (ESBS) and Senior Fellow at the Freiburg Institute for Advanced Studies. Biography Ralf Reski studied biology, chemistry and pedagogy at the Universities of Giessen and Hamburg. He was awarded his doctorate in Genetics in 1990 by the University of Hamburg and received his habilitation in General Botany in 1994. From 1996 until 1999, he was a Heisenberg-Fellow of the German Research Foundation. He was appointed Distinguished Professor and entitled Ordinarius at the University of Freiburg in 1999, where he became Head of the newly established Department of Plant Biotechnology. From 2001 until 2011, Reski was Director Plant Biotechnology at the Centre for Applied Biosciences (ZAB, University of Freiburg). Also since 2010 ...
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Fertilisation
Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Processes such as insemination or pollination which happen before the fusion of gametes are also sometimes informally called fertilisation. The cycle of fertilisation and development of new individuals is called sexual reproduction. During double fertilisation in angiosperms the haploid male gamete combines with two haploid polar nuclei to form a triploid primary endosperm nucleus by the process of vegetative fertilisation. History In Antiquity, Aristotle conceived the formation of new individuals through fusion of male and female fluids, with form and function emerging gradually, in a mode called by him as epigenetic. In 1784, Spallanzani established the need of interaction between the female's ovum and male's sperm to form a zygote in frog ...
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GUS Reporter System
The GUS reporter system (''GUS'': β-glucuronidase) is a reporter gene system, particularly useful in plant molecular biology and microbiology. Several kinds of GUS reporter gene assay are available, depending on the substrate used. The term GUS staining refers to the most common of these, a histochemical technique. Purpose The purpose of this technique is to analyze the activity of a gene transcription promoter (in terms of expression of a so-called reporter gene under the regulatory control of that promoter) either in a quantitative manner, involving some measure of activity, or qualitatively (on versus off) through visualization of its activity in different cells, tissues, or organs. The technique utilizes the uidA gene of Escherichia coli, which codes for the enzyme, β-glucuronidase; this enzyme, when incubated with specific colorless or non-fluorescent substrates, can convert them into stable colored or fluorescent products. The presence of the GUS-induced color indicates ...
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