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Archaeoistiodactylus
''Archaeoistiodactylus'' is an extinct genus of Wukongopteridae, wukongopterid pterosaur from the Middle Jurassic of China. Discovery and naming ''Archaeoistiodactylus'' is known from an incomplete skeleton with a partial skull and lower jaws. Catalogued as holotype specimen JPM04-0008, it was recovered from rocks of the Tiaojishan Formation in western Liaoning, China, which date to the Bathonian-Callovian stages of the Jurassic period. Apart from the skull, it preserves ribs, parts of the wings, hindlimbs and the pelvis. The neck, tail and feet of the specimen are lacking. ''Archaeoistiodactylus'' was first named and described by Lü Junchang and Fucha Xiaohui in 2010 in paleontology, 2010. The type (biology), type species is ''Archaeoistiodactylus linglongtaensis''. The generic name is a combination of the Greek ''archaios'', "ancient", with the name of the genus ''Istiodactylus'', referring to ''Archaeoistiodactylus'' being presumed to have been an older close relative of the ...
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Wukongopteridae
Wukongopteridae is a group of basal pterosaurs, found in China and the UK. It contains eight species in five genera, all dated to the Middle to Late Jurassic period, The Wukongopteridae were first named by Wang ''et al.'' in 2009, not yet giving an exact definition.* The clade Wukongopteridae was first defined by Wang ''et al.'' in 2010 as "the most recent common ancestor of '' Wukongopterus lii'' and '' Kunpengopterus sinensis'', and all of its descendants". Description Wukongopterids are characterized by a unique combination of "primitive" and advanced pterosaurian features. While they had long tails and other features characteristic of other "rhamphorhynchoids", they also had distinct pterodactyloid features, such as long vertebrae in the neck and a single skull opening in front of the eyes, the nasoantorbital fenestra (in most "rhamphorhynchoids", the antorbital fenestra and the nasal opening are separate). This feature lead to ''Darwinopterus modularis'' being placed by L ...
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Timeline Of Pterosaur Research
This timeline of pterosaur research is a chronologically ordered list of important fossil discoveries, controversies of interpretation, and taxonomic revisions of pterosaurs, the famed flying reptiles of the Mesozoic era. Although pterosaurs went extinct millions of years before humans evolved, humans have coexisted with pterosaur fossils for millennia. Before the development of paleontology as a formal science, these remains would have been interpreted through a mythological lens. Myths about thunderbirds told by the Native Americans of the modern Western United States may have been influenced by observations of ''Pteranodon'' fossils. These thunderbirds were said to have warred with water monsters, which agrees well with the co-occurrence of ''Pteranodon'' and the ancient marine reptiles of the seaway over which it flew. The formal study of pterosaurs began in the late 18th century when naturalist Cosimo Alessandro Collini of Mannheim, Germany published a description of ...
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Monofenestrata
Monofenestrata is an unranked group of pterosaurs that includes the family Wukongopteridae and the suborder Pterodactyloidea. The clade Monofenestrata was in 2009/2010 defined as the group consisting of ''Pterodactylus'' and all species sharing with ''Pterodactylus'' the synapomorphy of an external nostril confluent with the antorbital fenestra, the major skull opening on the side of the snout. The name is derived from Greek ''monos'', "single", and Latin ''fenestra'', "window". The concept was inspired by the discovery of ''Darwinopterus'', a species combining a pterodactyloid-type skull with a more basal build of the remainder of the body. The Darwinoptera, a primitive subgroup of monofenestratans showing this transitional anatomy, was also named for ''Darwinopterus'' and defined as all descendants of its common ancestor with '' Pterorhynchus''. The earliest known monofenestratan fossils have been found in the Stonesfield Slate formation of the United Kingdom, which dates to t ...
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Tiaojishan Formation
The Tiaojishan Formation is a geological formation in Hebei and Liaoning, People's Republic of China, dating to the middle-late Jurassic period (Bathonian- Oxfordian stages). It is known for its exceptionally preserved fossils, including those of plants, insects and vertebrates. It is made up mainly of pyroclastic rock interspersed with basic volcanic and sedimentary rocks. Previously, the Tiaojishan Formation was grouped together with the underlying Haifanggou Formation (also known as the Jiulongshan Formation) as a single "Lanqi Formation." The Tiaojishan Formation forms a key part of the Yanliao Biota assemblage, alongside the Haifanggou Formation. Age Using Argon–argon dating, Wang and colleagues in 2005 dated part of the Tiaojishan Formation to about 160 million years ago, the beginning of the Oxfordian stage, the first stage of the Upper Jurassic epoch. In 2006, a study by Liu and colleagues used U-Pb zircon dating to conclude that the Tiaojishan Formation correlates wi ...
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Istiodactylidae
Istiodactylidae is a small family of pterosaurs. This family was named in 2001 after the type genus ''Istiodactylus'' was discovered not to be a member of the genus ''Ornithodesmus''. Systematics and distribution Remains of taxa that can be confidently assigned to Istiodactylidae have been found in the UK and China, in rocks dating from the Early Cretaceous period (Barremian to Aptian stage). Arbour and Currie (2011) described Canadian '' Gwawinapterus beardi'' as a member of Istiodactylidae living in the late Cretaceous (upper Campanian stage); however, Witton (2012) suggested the tooth replacement pattern in this animal does not match that of pterosaurs, suggesting that the species might be non-pterosaurian. Additional research suggested that the species was in fact a fish. The earliest known species might be ''Archaeoistiodactylus linglongtaensis'', from the Middle Jurassic of China; however, it also has been suggested that the holotype specimen of this species might actuall ...
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Middle Jurassic
The Middle Jurassic is the second epoch of the Jurassic Period. It lasted from about 174.1 to 163.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relatively rare, but geological formations containing land animal fossils include the Forest Marble Formation in England, the Kilmaluag Formation in Scotland,British Geological Survey. 2011Stratigraphic framework for the Middle Jurassic strata of Great Britain and the adjoining continental shelf: research report RR/11/06 British Geological Survey, Keyworth, Nottingham. the Daohugou Beds in China, the Itat Formation in Russia, and the Isalo III Formation of western Madagascar. Paleogeography During the Middle Jurassic Epoch, Pangaea began to separate into Laurasia and Gondwana, and the Atlantic Ocean formed. Eastern Laurasia was tectonically active as the Cimmerian plate continued to collide with Laurasia's southern coast, completely closing the Paleo-Tethys Ocean. A subduction zone ...
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Istiodactylus
''Istiodactylus'' is a genus of pterosaur that lived during the Early Cretaceous period, about 120 million years ago. The first fossil was discovered on the English Isle of Wight in 1887, and in 1901 became the holotype specimen of a new species, ''O. latidens'' (Latin for "wide tooth"), in the genus '' Ornithodesmus''. This species was moved to its own genus, ''Istiodactylus'', in 2001; this name is Greek for "sail finger". More specimens were described in 1913, and ''Istiodactylus'' was the only pterosaur known from three-dimensionally preserved fossils for much of the 20th century. In 2006, a species from China, ''I. sinensis'', was assigned to ''Istiodactylus'', but it has also been suggested to belong to a different genus. ''Istiodactylus'' was a large pterosaur; estimates of its wingspan range from . Its skull was about long, and was relatively short and broad for a pterosaur. The front of the snout was low and blunt, and bore a semicircle of 48 teeth. The triangular teeth ...
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Middle Jurassic Pterosaurs Of Asia
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Kunpengopterus
''Kunpengopterus'' is a genus of wukongopterid pterosaur from the middle-late Jurassic Tiaojishan Formation of northeastern China. The genus contains two species, the type species ''K. sinensis'' and ''K. antipollicatus''. History of discovery ''K. sinensis'' is known from the holotype specimen IVPP V16047, an almost complete skeleton with complete skull and lower jaws recovered from rocks of the Tiaojishan Formation or Daohugou Beds in Linglongta, Jianchang County, western Liaoning. The age of these layers is controversial. This compression fossil is of an adult individual. Aside from the bones some soft parts were also preserved and the remains of a possibly regurgitated fish. ''Kunpengopterus'' was named and described by Wang Xiaolin, Alexander Wilhelm Armin Kellner, Jiang Shunxing, Cheng Xin, Meng Xi and Taissa Rodrigues in 2010. The type species is ''Kunpengopterus sinensis''. The generic name combines the Kun, a large fish or whale from Chinese folklore that could t ...
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Polytomy
An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree that contains any multifurcations can be described as a multifurcating tree. Soft polytomies vs. hard polytomies Two types of polytomies are recognised, soft and hard polytomies. Soft polytomies are the result of insufficient phylogenetic information: though the lineages diverged at different times – meaning that some of these lineages are closer relatives than others, and the available data does not allow recognition of this. Most polytomies are soft, meaning that they would be resolved into a typical tree of dichotomies if better data were available. In contrast, a hard polytomy represents a true divergence event of three or more lineages. Applications Interpretations for a polytomy depend on the individuals, that are represe ...
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Darwinopterus
''Darwinopterus'' (meaning "Darwin's wing") is a genus of pterosaur, discovered in China and named after biologist Charles Darwin. Between 30 and 40 fossil specimens have been identified, all collected from the Tiaojishan Formation, which dates to the middle Jurassic period, 160.89–160.25 Ma ago.Liu Y.-Q. Kuang H.-W., Jiang X.-J., Peng N., Xu H. & Sun H.-Y. (2012). "Timing of the earliest known feathered dinosaurs and transitional pterosaurs older than the Jehol Biota." ''Palaeogeography, Palaeoclimatology, Palaeoecology'' (advance online publication). The type species, type species, ''D. modularis'', was described in February 2010 in paleontology, 2010. ''D. modularis'' was the first known pterosaur to display features of both long-tailed (Rhamphorhynchoidea, rhamphorhynchoid) and short-tailed (Pterodactyloidea, pterodactyloid) pterosaurs, and was described as a transitional fossil between the two groups.Dell'Amore, C. (2009)"Odd New Pterosaur: 'Darwin's Wing' Fills Evolution Ga ...
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