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Anteophthalmosuchus
''Anteophthalmosuchus'' (meaning "forward-pointing eye crocodile") is an extinct genus of goniopholidid mesoeucrocodylian from the Early Cretaceous of southern England, eastern Spain, and western Belgium. Discovery The holotype specimen of ''Anteophthalmosuchus'', from the Wealden Group of the Isle of Wight, includes a well-preserved skull and partial skeleton. This specimen has been known since 1904 and was identified as the "Tie Pits specimen" or the "Hooley specimen" after Reginald Walter Hooley, an amateur paleontologist who had described it in 1905. Hooley had originally attributed the specimen to the previously named species '' Goniopholis crassidens''. Additional referred specimens include a partial disarticulated skeleton and a partial skull that may represent a juvenile specimen. In 2011, Hooley's specimen was redescribed as a distinct genus and species of goniopholidid called ''Anteophthalmosuchus hooleyi''. The genus name means "forward-pointing eye crocodile" becaus ...
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Anteophthalmosuchus
''Anteophthalmosuchus'' (meaning "forward-pointing eye crocodile") is an extinct genus of goniopholidid mesoeucrocodylian from the Early Cretaceous of southern England, eastern Spain, and western Belgium. Discovery The holotype specimen of ''Anteophthalmosuchus'', from the Wealden Group of the Isle of Wight, includes a well-preserved skull and partial skeleton. This specimen has been known since 1904 and was identified as the "Tie Pits specimen" or the "Hooley specimen" after Reginald Walter Hooley, an amateur paleontologist who had described it in 1905. Hooley had originally attributed the specimen to the previously named species '' Goniopholis crassidens''. Additional referred specimens include a partial disarticulated skeleton and a partial skull that may represent a juvenile specimen. In 2011, Hooley's specimen was redescribed as a distinct genus and species of goniopholidid called ''Anteophthalmosuchus hooleyi''. The genus name means "forward-pointing eye crocodile" becaus ...
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Hulkepholis Plotos
''Hulkepholis'' is an extinct genus of goniopholidid mesoeucrocodylian from the Early Cretaceous of southern England and eastern Spain. It contains two species, the type species, ''Hulkepholis willetti'', and also ''H. plotos''. ''Hulkepholis'' is most closely related to both species of ''Anteophthalmosuchus'' (including "Dollo's goniopholidid"). Discovery ''H. willetti'' is known from a single holotype specimen, BMNHB 001876 from the Wealden Group of the Isle of Wight that includes a well-preserved nearly complete skull. It was collected by Edgar W. Willett at Cuckfield, West Sussex, from the Valanginian-aged Grinstead Clay Member, of the Hastings Group, Wealden Supergroup. Willett showed the specimen to the Geological Society of London in or around 1877 or 1878, and it was then Hulke (1878) described it as a specimen of ''Goniopholis crassidens'', an identification derived from a similar tooth form. The specimen was considered lost among the crocodyliform research community bu ...
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Goniopholis Crassidens
''Goniopholis'' (meaning "angled scale") is an extinct genus of goniopholidid crocodyliform that lived in Europe and Africa during the Late Jurassic and Early Cretaceous. Being semi-aquatic it is very similar to modern crocodiles. It ranged from 2–4 metres in length, and would have had a very similar lifestyle to the American alligator or Nile crocodile. Discovery and species The type species of the genus is ''G. crassidens'' which is known from the Berriasian of England, and the referable species ''G simus'' from the Berriasian of NW Germany, might be conspecific. Other species that are referable to ''Goniopholis'' include ''G. kiplingi'' from the Berriasian of England, and ''G. baryglyphaeus'' from the Late Jurassic (Kimmeridgian) of Portugal making it the oldest known ''Goniopholis'' species. The species ''G. kiplingi'' honors the author Rudyard Kipling, "in recognition for his enthusiasm for natural sciences". Eggs attributed to ''Goniopholis'' were found in the Late Jura ...
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Goniopholidid
Goniopholididae is an extinct family of moderate-sized semi-aquatic neosuchian crocodyliformes. Their bodyplan and morphology are convergent on living crocodilians. They lived across Laurasia (Asia, Europe and North America) between the Middle Jurassic (possibly Early Jurassic, see below) and the Late Cretaceous. Description Compared to modern crocodilians, goniopholidids are very unusual in several respects. They possessed two rows of rectangular, interlocking osteoderms like those of terrestrial crocodilymorphs like atoposaurids, that are relatively simple, do not extend far in their necks, as opposed to the ornate armours of modern crocodilians; likewise, unlike modern crocodilians but like many extinct forms like phytosaurs, they have ventral osteoderms as well. Their forelimbs are also proportionally very long, particularly in the humeri and wrist bones, being as long or longer than the hindlimbs, the opposite of the condition seen in modern crocodilians. Some like ''Anteoph ...
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Goniopholis Simus 1
''Goniopholis'' (meaning "angled scale") is an extinct genus of goniopholidid crocodyliform that lived in Europe and Africa during the Late Jurassic and Early Cretaceous. Being semi-aquatic it is very similar to modern crocodiles. It ranged from 2–4 metres in length, and would have had a very similar lifestyle to the American alligator or Nile crocodile. Discovery and species The type species of the genus is ''G. crassidens'' which is known from the Berriasian of England, and the referable species ''G simus'' from the Berriasian of NW Germany, might be conspecific. Other species that are referable to ''Goniopholis'' include ''G. kiplingi'' from the Berriasian of England, and ''G. baryglyphaeus'' from the Late Jurassic (Kimmeridgian) of Portugal making it the oldest known ''Goniopholis'' species. The species ''G. kiplingi'' honors the author Rudyard Kipling, "in recognition for his enthusiasm for natural sciences". Eggs attributed to ''Goniopholis'' were found in the Late Jura ...
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Sainte-Barbe Clays Formation
The Sainte-Barbe Clays Formation is a geological formation in Belgium. It is found in localised areas of the northern margin of the Mons Basin, alongside the equivalently aged Hautrage and Baudour Clay Formations. It is Upper Barremian-Lower Aptian in age. It predominantly consists of laminated clay, with some lignite. It is well known for the "Iguanodon sinkhole" locality near Bernissart where many specimens of Iguanodon, ''Iguanodon bernissartensis'' were described by Louis Dollo in the late 19th century. Geologic context The Sainte-Barbe Clays Formation are part of the stratigraphy of the Mons Basin, a Cretaceous-Cenozoic sedimentary basin in western Belgium. The subsidence of the basin floor is caused by the Solvation, dissolution of anhydrite in the Carboniferous Basement (geology), basement. Wealden facies are the oldest units within the basin, and are only found on the northern margin of the basin, being absent from the central and southern parts of the basin. Localised are ...
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Escucha Formation
The Escucha Formation is a geological formation in La Rioja and Teruel provinces of northeastern Spain whose strata date back to the late Aptian to middle Albian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, et al., 2004 The approximately thick formation underlies the Utrillas Formation and overlies Castrillo de la Reina, Benassal & Oliete Formations. The Escucha Formation comprises siltstones, mudstones, sandstones, coal, siltstones and amber, in which several fossil insects were found. The formation was deposited in a variety of continental to paralic (deltaic) environments.Peyrot et al., 2007 Fossil content The Escucha Formation has provided the following fossils, among others:
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Early Cretaceous
The Early Cretaceous ( geochronological name) or the Lower Cretaceous (chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145  Ma to 100.5 Ma. Geology Proposals for the exact age of the Barremian-Aptian boundary ranged from 126 to 117 Ma until recently (as of 2019), but based on drillholes in Svalbard the defining early Aptian Oceanic Anoxic Event 1a (OAE1a) was carbon isotope dated to 123.1±0.3 Ma, limiting the possible range for the boundary to c. 122–121 Ma. There is a possible link between this anoxic event and a series of Early Cretaceous large igneous provinces (LIP). The Ontong Java-Manihiki-Hikurangi large igneous province, emplaced in the South Pacific at c. 120 Ma, is by far the largest LIP in Earth's history. The Ontong Java Plateau today covers an area of 1,860,000 km2. In the Indian Ocean another LIP began to form at c. 120 Ma, the Kerguelen P ...
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Frontal Bone
The frontal bone is a bone in the human skull. The bone consists of two portions.''Gray's Anatomy'' (1918) These are the vertically oriented squamous part, and the horizontally oriented orbital part, making up the bony part of the forehead, part of the bony orbital cavity holding the eye, and part of the bony part of the nose respectively. The name comes from the Latin word ''frons'' (meaning " forehead"). Structure of the frontal bone The frontal bone is made up of two main parts. These are the squamous part, and the orbital part. The squamous part marks the vertical, flat, and also the biggest part, and the main region of the forehead. The orbital part is the horizontal and second biggest region of the frontal bone. It enters into the formation of the roofs of the orbital and nasal cavities. Sometimes a third part is included as the nasal part of the frontal bone, and sometimes this is included with the squamous part. The nasal part is between the brow ridges, and ends in ...
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Anatomical Terms Of Location
Standard anatomical terms of location are used to unambiguously describe the anatomy of animals, including humans. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position provides a definition of what is at the front ("anterior"), behind ("posterior") and so on. As part of defining and describing terms, the body is described through the use of anatomical planes and anatomical axes. The meaning of terms that are used can change depending on whether an organism is bipedal or quadrupedal. Additionally, for some animals such as invertebrates, some terms may not have any meaning at all; for example, an animal that is radially symmetrical will have no anterior surface, but can still have a description that a part is close to the middle ("proximal") or further from the middle ("distal"). International organisations have determined vocabularies that are often used as standard vocabularies for subdisciplines of anatom ...
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Holotype
A holotype is a single physical example (or illustration) of an organism, known to have been used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of several examples, but explicitly designated as the holotype. Under the International Code of Zoological Nomenclature (ICZN), a holotype is one of several kinds of name-bearing types. In the International Code of Nomenclature for algae, fungi, and plants (ICN) and ICZN, the definitions of types are similar in intent but not identical in terminology or underlying concept. For example, the holotype for the butterfly '' Plebejus idas longinus'' is a preserved specimen of that subspecies, held by the Museum of Comparative Zoology at Harvard University. In botany, an isotype is a duplicate of the holotype, where holotype and isotypes are often pieces from the same individual plant or samples from the same gathering. A holotype is not necessarily "typ ...
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Occipital Condyle
The occipital condyles are undersurface protuberances of the occipital bone in vertebrates, which function in articulation with the superior facets of the atlas vertebra. The condyles are oval or reniform (kidney-shaped) in shape, and their anterior extremities, directed forward and medialward, are closer together than their posterior, and encroach on the basilar portion of the bone; the posterior extremities extend back to the level of the middle of the foramen magnum. The articular surfaces of the condyles are convex from before backward and from side to side, and look downward and lateralward. To their margins are attached the capsules of the atlanto-occipital joints, and on the medial side of each is a rough impression or tubercle for the alar ligament. At the base of either condyle the bone is tunnelled by a short canal, the hypoglossal canal. Clinical significance Fracture of an occipital condyle may occur in isolation, or as part of a more extended basilar skull fracture ...
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