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Anomocephaloidea
Anomocephaloidea is a clade (evolutionary grouping) of anomodont therapsids that existed in Gondwana during the Middle Permian and includes two species, ''Anomocephalus africanus'' from South Africa and ''Tiarajudens eccentricus'' from Brazil, both of which are characterized by large body size and teeth that fit tightly together or occlude. Anomocephaloidea is among the most basal groups of anomodonts, the other being Venyukovioidea, which differs in being a Laurasian clade of mostly small-bodied species. Anomocephaloidea was named in 2011 with the discovery of ''Tiarajudens''; ''Anomocephalus'' had been known since 1999, but was unique among anomodonts until ''Tiarajudens'' was described. Both ''Anomocephalus'' and ''Tiarajudens'' were herbivores, although the latter possessed a pair of saber-like canine teeth that may have been used in display or combat with other individuals of the same species. Although Anomocephaloidea was short-lived and had a limited geographic distribut ...
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Venyukovioidea
Venyukovioidea is an infraorder of anomodont therapsids related to dicynodonts from the Permian of Russia. They have also known as Venjukovioidea, as well as by the similar names Venyukoviamorpha or Venjukoviamorpha in literature. This in part owes to a misspelling by Russian palaeontologist Ivan Efremov in 1940 when he mistakenly spelt '' Venyukovia'', the namesake of the group, with a 'j' instead of a 'y' (i.e. ''Venjukovia''), which permeated through subsequent therapsid literature before the mistake was caught and corrected. The order Ulemicia has also been coined for a similar taxonomic concept in Russian scientific literature, which notably excludes ''Suminia'' and ''Parasuminia''. Venyukovioidea includes the genera ''Venyukovia'', '' Otsheria'', '' Ulemica'', '' Suminia'' and ''Parasuminia'', all from Western Siberia. Historically, some of these genera have been placed in various families and subfamilies, including the Venyukoviidae/Venjukoviidae, Otsheriidae, and Ule ...
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Anomocephalus Africanus
''Anomocephalus'' is an extinct genus of primitive anomodonts and belongs to the clade Anomocephaloidea. The name is said to be derived from the Greek word ''anomos'' meaning lawless and ''cephalos'' meaning head. The proper word for head in Greek is however κεφαλή (''kephalē'').Liddell, H.G. & Scott, R. (1940). ''A Greek-English Lexicon. revised and augmented throughout by Sir Henry Stuart Jones. with the assistance of. Roderick McKenzie.'' Oxford: Clarendon Press. It is primitive in that it retains a complete set of teeth in both jaws, in contrast to its descendants, the dicynodonts, whose dentition is reduced to only a single pair of tusks (and in many cases no teeth at all), with their jaws covered by a horny beak similar to that of a modern tortoise. However, they are in no way closely related. Its discovery in 1999 from the earliest terrestrial rocks of Gondwana (from Williston in the Karoo of the Northern Cape Province of South Africa) has shown that this grou ...
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Tiarajudens Eccentricus
''Tiarajudens'' (" Tiaraju tooth") is an extinct genus of saber-toothed herbivorous anomodonts which lived during the Middle Permian period (Capitanian stage) in what is now Rio Grande do Sul, Brazil. It is known from the holotype UFRGS PV393P, a nearly complete skull. The type species ''T. eccentricus'' was named in 2011. Description ''Tiarajudens'' is a member of Anomodontia, a suborder of therapsids. Like other anomodonts, it was a quadrupedal herbivore about the size of a wild boar. The single fragmented holotype skull is short and robust at about in length. The most prominent features of ''Tiarajudens'' are its two large saber-like canine teeth. These teeth are unlike the tusks of dicynodonts, a later group of anomodonts. Twenty-one high- crowned teeth are present on either side of the upper jaw, including spoon-shaped incisors. Wide palatal teeth are also present. The top and bottom sets of teeth fit closely together, much like the teeth of mammals, allowing i ...
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Anomocephalus
''Anomocephalus'' is an extinct genus of primitive anomodonts and belongs to the clade Anomocephaloidea. The name is said to be derived from the Greek word ''anomos'' meaning lawless and ''cephalos'' meaning head. The proper word for head in Greek is however κεφαλή (''kephalē'').Liddell, H.G. & Scott, R. (1940). ''A Greek-English Lexicon. revised and augmented throughout by Sir Henry Stuart Jones. with the assistance of. Roderick McKenzie.'' Oxford: Clarendon Press. It is primitive in that it retains a complete set of teeth in both jaws, in contrast to its descendants, the dicynodonts, whose dentition is reduced to only a single pair of tusks (and in many cases no teeth at all), with their jaws covered by a horny beak similar to that of a modern tortoise. However, they are in no way closely related. Its discovery in 1999 from the earliest terrestrial rocks of Gondwana (from Williston in the Karoo of the Northern Cape Province of South Africa) has shown that this group ...
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Tiarajudens
''Tiarajudens'' (" Tiaraju tooth") is an extinct genus of saber-toothed herbivorous anomodonts which lived during the Middle Permian period ( Capitanian stage) in what is now Rio Grande do Sul, Brazil. It is known from the holotype UFRGS PV393P, a nearly complete skull. The type species ''T. eccentricus'' was named in 2011. Description ''Tiarajudens'' is a member of Anomodontia, a suborder of therapsids. Like other anomodonts, it was a quadrupedal herbivore about the size of a wild boar. The single fragmented holotype skull is short and robust at about in length. The most prominent features of ''Tiarajudens'' are its two large saber-like canine teeth. These teeth are unlike the tusks of dicynodonts, a later group of anomodonts. Twenty-one high- crowned teeth are present on either side of the upper jaw, including spoon-shaped incisors. Wide palatal teeth are also present. The top and bottom sets of teeth fit closely together, much like the teeth of mammals, allowing it t ...
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Anomodont
Anomodontia is an extinct group of non-mammalian therapsids from the Permian and Triassic periods. By far the most speciose group are the dicynodonts, a clade of beaked, tusked herbivores.Chinsamy-Turan, A. (2011) ''Forerunners of Mammals: Radiation - Histology - Biology''p.39.Indiana University Press, . Retrieved May 2012 Anomodonts were very diverse during the Middle Permian, including primitive forms like ''Anomocephalus'' and ''Patranomodon'' and groups like Venyukovioidea and Dromasauria. Dicynodonts became the most successful and abundant of all herbivores in the Late Permian, filling ecological niches ranging from large browsers down to small burrowers. Few dicynodont families survived the Permian–Triassic extinction event, but one lineage (Kannemeyeriiformes) evolved into large, stocky forms that became dominant terrestrial herbivores right until the Late Triassic, when changing conditions caused them to decline, finally going extinct during the Triassic-Jurassic ext ...
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Tetraceratops
''Tetraceratops insignis'' ("four-horned face emblem") is an extinct synapsid from the Early Permian that was formerly considered the earliest known representative of Therapsida, a group that includes mammals and their close extinct relatives. It is known from a single skull, discovered in Texas in 1908. According to a 2020 study, it should be classified as a primitive non-therapsid sphenacodont rather than a genuine basal therapsid. Description ''Tetraceratops'' is known from a single skull discovered in Texas in the early 1900s. Contrary to its genus name, ''Tetraceratops'' actually has six horns, one pair being on the premaxilla bones, one pair on the prefrontal bones, and one pair on the angular processes of the mandible. When it was discovered and described in 1908, the skull was still embedded in a matrix, and only the premaxilla and prefrontal pairs were visible. In life, thus, it would have resembled a large lizard with four horns on its snout, and a pair of large s ...
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Raranimus
''Raranimus'' is an extinct genus of therapsids of the Middle Permian. It was described in 2009 from a partial skull found in 1998 from the Dashankou locality of the Xidagou Formation, outcropping in the Qilian Mountains of Gansu, China. The genus is the most basal known member of the clade Therapsida, to which the later Mammalia belong. Description ''Raranimus'' shares a number of features with later therapsids and ancestral Sphenacodontia. The skull consists of a well preserved rostrum. The teeth suggest a carnivorous lifestyle for ''Raranimus'', as the incisors are recurved and the second canines are serrated on their posterior edges. The incisors are morphologically similar to those seen in more derived theriodonts. The presence of two linguo−labially compressed canines is a diagnostic feature of ''Raranimus''. The presence of two functional canines is characteristic of sphenacodontids, and this condition is seen in no other therapsid other than ''Rananimus''. However, th ...
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Lycosuchus
''Lycosuchus'' ("wolf crocodile") is an extinct genus of carnivorous therocephalians which lived in the Middle Permian 265—260 Ma existing for approximately . As a member of the Lycosuchidae, the genus represents one of the earliest diverging therocephalians. The type species ''Lycosuchus vanderrieti'' is known from a handful of well-preserved specimens featuring the cranium and lower jaw; the holotype US D173 housed at the University of Stellenbosch, South Africa, is a near complete occluded skull. Specimen MB.R. 995, housed at the Museum für Naturkunde Berlin, Germany, consists of a near complete and isolated lower jaw, along with a partial snout and brain case. With the help of μCT data, Pusch et al (2020) described the endocranial anatomy of ''Lycosuchus vanderrieti''. It was a medium-sized predator, reaching 1.2 m (3.8 ft) in length with a skull 23 cm long., typical of early therocephalians. ''L. vanderrieti'' bore two functional canines in each maxil ...
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Biarmosuchus
''Biarmosuchus'' is an extinct genus of biarmosuchian therapsids that lived around 267 mya during the Middle Permian period. ''Biarmosuchus'' was discovered in the Perm region of Russia. The first specimen was found in channel sandstone that was deposited by flood waters originating from the young Ural Mountains. Description ''Biarmosuchus'' was a medium-sized predator, similar in size to a large dog, grew up to 1.5–2 m in length with a skull length 15 cm (immature) to 21 cm. It was a lightly built, probably agile animal that would have fed on smaller tetrapods. Their legs are quite long, and the animals were probably quite agile in spite of their size. A large opening for the eye and a small temple opening common in primitive stem-mammals, this lends to a weak bite but how it ate is pure speculation. The teeth contained eight small incisors on the palate, followed by a canine tooth and a further five canine teeth. So together the species contained fourteen uppe ...
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Middle Permian
The Guadalupian is the second and middle series/ epoch of the Permian. The Guadalupian was preceded by the Cisuralian and followed by the Lopingian. It is named after the Guadalupe Mountains of New Mexico and Texas, and dates between 272.95 ± 0.5 – 259.1 ± 0.4 Mya. The series saw the rise of the therapsids, a minor extinction event called Olson's Extinction and a significant mass extinction called the end-Capitanian extinction event. The Guadalupian was previously known as the Middle Permian. Name and background The Guadalupian is the second and middle series or epoch of the Permian. Previously called Middle Permian, the name of this epoch is part of a revision of Permian stratigraphy for standard global correlation. The name "Guadalupian" was first proposed in the early 1900s, and approved by the International Subcommission on Permian Stratigraphy in 1996. References to the Middle Permian still exist. The Guadalupian was preceded by the Cisuralian and followed by t ...
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Gorgonops
''Gorgonops'' (from el, Γοργών 'Gorgon' and 'eye, face', literally 'Gorgon eye' or 'Gorgon face') is an extinct genus of gorgonopsian therapsids, of which it is the type genus, having lived during the Late Permian (Wuchiapingian), about 260–254 million years ago in what is now South Africa. Despite its popularity, ''Gorgonops'' is a medium-sized gorgonopsian (about long maximum), regularly confused by the general public with the more massive '' Inostrancevia'', known from Russia, due to their similar appearance and the various media that tend to refer them by the name of the group they belong rather than by their genus names, which does not help in differentiation. History of discovery The holotype of the type species, ''Gorgonops torvus'', was in 1876 one of the first therapsids described, by Richard Owen, who also coined the name "Dinosauria" on the basis of the first known dinosaur fossils. It was also used as the type for which Richard Lydekker described the famil ...
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