Ankylosaur
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Ankylosaur
Ankylosauria is a group of herbivorous dinosaurs of the order Ornithischia. It includes the great majority of dinosaurs with armor in the form of bony osteoderms, similar to turtles. Ankylosaurs were bulky quadrupeds, with short, powerful limbs. They are known to have first appeared in the Middle Jurassic, and persisted until the end of the Cretaceous Period. The two main families of Ankylosaurs, Nodosauridae and Ankylosauridae are primarily known from the Northern Hemisphere, but the more basal Parankylosauria are known from southern Gondwana during the Cretaceous. Ankylosauria was first named by Henry Fairfield Osborn in 1923.Osborn, H. F. (1923). "Two Lower Cretaceous dinosaurs of Mongolia." ''American Museum Novitates'', 95: 1–1/ref> In the Linnaean classification system, the group is usually considered either a suborder or an infraorder. It is contained within the group Thyreophora, which also includes the stegosaurs, armored dinosaurs known for their combination of plate ...
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Mymoorapelta
''Mymoorapelta'' (Meaning "Vannetta Moore and Pete and Marilyn Mygatt's shield" after a combination of the names of the discoverers of the Mygatt-Moore Quarry that fossils were originally collected from, and pɛltə "shield") is a monospecific genus of nodosaurid ankylosaur, a group of heavily armored, herbivorous, quadrupedal dinosaurs, from the Late Jurassic (Kimmeridgian- Tithonian, around 155 to 150 million years ago) Morrison Formation (Brushy Basin Member) of western Colorado and central Utah, USA. Few specimens are known, but the most complete one is the holotype individual from the Mygatt-Moore Quarry that includes many osteoderms, a partial skull, vertebrae, and other bones. It was initially described by James Kirkland and Kenneth Carpenter in 1994. Along with ''Gargoyleosaurus'', it is one of the earliest known nodosaurids. ''Mymoorapleta'' is one of the smaller known nodosaurids, with the estimated length of the largest specimen only reaching . It had a narrow snou ...
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Parankylosauria
Parankylosauria is a group of basal ankylosaurian dinosaurs known from the Cretaceous of South America, Antarctica, and Australia. It is thought the group split from other ankylosaurs during the mid-Jurassic period, despite this being unpreserved in the fossil record. History of research During the Mesozoic era, the southern continents (South America, Antarctica, Australia, and Africa in addition to India and Zealandia) were unified into a supercontinent known as Gondwana. This was in contrast to the supercontinent of Laurasia in the Northern Hemisphere, with both originating from the breakup of Pangaea. Gondwana itself gradually split apart over the course of the Jurassic and Cretaceous eras. Ankylosaurian dinosaurs from Laurasia have historically been far more extensively recorded and studied. Reports of the group in Gondwana date back to 1904, with a specimen from Australia and include referrals of ''Loricosaurus'', ''Lametasaurus'', and ''Brachypodosaurus'' to group among ass ...
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Ankylosauridae
Ankylosauridae () is a family of armored dinosaurs within Ankylosauria, and is the sister group to Nodosauridae. The oldest known Ankylosaurids date to around 122 million years ago and went extinct 66 million years ago during the Cretaceous–Paleogene extinction event. These animals were mainly herbivorous and were obligate quadrupeds, with leaf-shaped teeth and robust, scute-covered bodies. Ankylosaurids possess a distinctly domed and short snout, wedge-shaped osteoderms on their skull, scutes along their torso, and a tail club. Ankylosauridae is exclusively known from the northern hemisphere, with specimens found in western North America, Europe, and East Asia. The first discoveries within this family were of the genus ''Ankylosaurus'', by Peter Kaiser and Barnum Brown in Montana in 1906. Brown went on to name Ankylosauridae and the subfamily Ankylosaurinae in 1908. Anatomy Ankylosaurids are stout, solidly built, armoured dinosaurs. They possess accessory ossifications on c ...
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Akainacephalus
''Akainacephalus'' (meaning "thorn head") is a monospecific genus of ankylosaurid dinosaur from southern Utah that lived during the Late Cretaceous (late Campanian, 76.26 Ma) in what is now the Horse Mountain Gryposaur Quarry of the Kaiparowits Formation. The type and only species, ''Akainacephalus johnsoni'', is known from the most complete ankylosaur specimen ever discovered from southern Laramidia, including a complete skull, tail club, a number of osteoderms, limb elements and part of its pelvis, among other remains. It was described in 2018 by Jelle P. Wiersma and Randall B. Irmis. It is closely related and shares similar cranial anatomy to ''Nodocephalosaurus''. Discovery and naming An almost complete skeleton of an ankylosaurid was excavated during the 2008, 2009, and 2010 field seasons from the Horse Mountain Gryposaur (HMG) Quarry in the Grand Staircase–Escalante National Monument, Kane County, Utah. The Horse Mountain Gryposaur Quarry represents a multitaxic and mult ...
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Tianzhenosaurus
''Tianzhenosaurus'' (meaning “ Tianzhen lizard”) is a monospecific genus of ankylosaurid dinosaur from the Shanxi Province that lived during the Late Cretaceous (Cenomanian-Campanian, ~99-71 Ma) in what is now the Huiquanpu Formation. ''Tianzhenosaurus'' may represent a junior synonym of ''Saichania'', an ankylosaurine known from the Barun Goyot and Nemegt Formation. Discovery and naming In 1983, Pang Qiqing and Cheng Zhengwu discovered articulated cervical vertebrae of an ankylosaurid from the Shanxi Province. Numerous excavations at the site yielded more than 2,300 specimens belonging to sauropods, theropods, ornithopods and ankylosaurid specimens. The holotype specimen, HBV-10001, consists of a partial skull. Two paratype specimens were assigned to ''Tianzhenosaurus'': HBV-10002, an incomplete mandible; HBV-10003, cervical vertebrae, dorsal vertebrae, caudal vertebrae, a sacral complex, ilia, pectoral girdles, pelvic girdles, fore and hind limbs with fore and hind feet, ta ...
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Nodosauridae
Nodosauridae is a family of ankylosaurian dinosaurs, from the Late Jurassic to the Late Cretaceous period in what is now North America, South America, Europe, and Asia. Description Nodosaurids, like their close relatives the ankylosaurids, were heavily armored dinosaurs adorned with rows of bony armor nodules and spines (osteoderms), which were covered in keratin sheaths. All nodosaurids, like other ankylosaurians, were medium-sized to large, heavily built, quadrupedal, herbivorous dinosaurs, possessing small, leaf-shaped teeth. Unlike ankylosaurids, nodosaurids lacked mace-like tail clubs, instead having flexible tail tips. Many nodosaurids had spikes projecting outward from their shoulders. One particularly well-preserved nodosaurid "mummy", known as the Suncor nodosaur (''Borealopelta markmitchelli''), preserved a nearly complete set of armor in life position, as well as the keratin covering and mineralized remains of the underlying skin, which indicate reddish dorsal pigmen ...
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Dracopelta
''Dracopelta'' (meaning “dragon shield”) is a monospecific genus of ankylosaur dinosaur from Portugal that lived during the Late Jurassic (uppermost lower Tithonian-upper Tithonian, 152.1-145.0 Ma) in what is now the Lourinhã Formation. The type and only species is ''Dracopelta zbyszewskii'', which is represented by a partial skeleton including unpublished material. Discovery and naming In 1963 or, more likely, early 1964, a partial skeleton was discovered during road construction works between the village of Barril and Praia da Assenta. Leonel Trindade confirmed the presence of the fossil in the area and photographed the specimen ''in situ''. Georges Zbyszewski and Octávio da Veiga Ferreir organized the excavation and extraction of the specimen in December 1964. Parts of the specimen were unprepared and misplaced, mixed in with a specimen of ''Miragaia''. The holotype specimen, MG 3 (formerly IGM 5787 and IGM 3), consists of dorsal vertebrae, articulated proximal ribs, o ...
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Liaoningosaurus
''Liaoningosaurus'' is an unusual genus of ankylosaurian dinosaurs from the Early Cretaceous period of China. It contains a single species, ''Liaoningosaurus paradoxus'', and is represented by two fossil specimens collected from the Yixian Formation (Aptian age) of Liaoning. ''L. paradoxus'' was unusual among advanced ornithischian dinosaurs in that it is speculated to have hunted or scavenged, with preserved gut contents showing that it may have eaten fish. Additionally, some features of its skeleton may suggest that it was partially aquatic.Ji Q., Wu X., Cheng Y., Ten F., Wang X., and Ji Y. 2016Fish-hunting ankylosaurs (Dinosauria, Ornithischia) from the Cretaceous of China ''Journal of Geology'', 40(2) . The type species ''L. paradoxus'' was named in 2001. The generic name refers to Liaoning. The specific name refers to the confusing mix of nodosaurid and ankylosaurid features shown by the specimen. Discovery The holotype IVPP V12560 is an articulated skeleton measuring ...
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Dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is the subject of active research. They became the dominant terrestrial vertebrates after the Triassic–Jurassic extinction event 201.3 mya; their dominance continued throughout the Jurassic and Cretaceous periods. The fossil record shows that birds are feathered dinosaurs, having evolved from earlier theropods during the Late Jurassic epoch, and are the only dinosaur lineage known to have survived the Cretaceous–Paleogene extinction event approximately 66 mya. Dinosaurs can therefore be divided into avian dinosaurs—birds—and the extinct non-avian dinosaurs, which are all dinosaurs other than birds. Dinosaurs are varied from taxonomic, morphological and ecological standpoints. Birds, at over 10,700 living species, are among ...
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Gastonia (dinosaur)
''Gastonia'' is a genus of herbivorous ankylosaurian dinosaur from the Early Cretaceous of North America, around 139 to 125 million years ago. It is often considered a nodosaurid closely related to ''Polacanthus''. ''Gastonia'' has a sacral shield and large shoulder spikes. Discovery and species The type specimen of ''Gastonia burgei'' ( CEUM 1307) was discovered in a bonebed from the limestone strata of the lower Cedar Mountain Formation in Yellow Cat Quarry, Grand County, eastern Utah, the type specimen consisting of a single skull. The type specimen was found alongside 4 partial skeletons of ''Gastonia that'' were placed as paratypes, along with the type specimen of ''Utahraptor'' and an Iguanodontid. ''Gastonia'' is among the most common dinosaur fossils in the Cedar Mountain Formation, with many individuals being found across several quarries in the southwest.Kirkland, J.I. (1998). A polacanthine ankylosaur (Ornithischia: Dinosauria) from the Early Cretaceous (Barremian) o ...
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Scolosaurus
''Scolosaurus'' is an extinct genus of ankylosaurid dinosaurs within the subfamily Ankylosaurinae. It is known from the lower levels of the Dinosaur Park Formation and upper levels of the Oldman Formation in the Late Cretaceous (latest middle Campanian stage, about 76.5 Ma ago) of Alberta, Canada. It contains two species, ''S. cutleri'' and ''S. thronus''. The type species, ''S. cutleri'', measured up to in length and in body mass. Discovery ''Scolosaurus'' was named by Franz Nopcsa von Felső-Szilvás in 1928, based on holotype NHMUK R.5161, a nearly complete specimen that preserves the entire skeleton except for the distal end of the tail, the right forelimb, the right hindlimb, and the skull. The rare preservation of osteoderms and skin impression are also present. The fossil skeleton was discovered by William Edmund Cutler, an independent fossil collector in 1914 at Quarry 80 of the Deadlodge Canyon locality. It was collected from the bottom of the Dinosaur Park ...
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Edmontonia
''Edmontonia'' is a genus of panoplosaurin nodosaurid dinosaur from the Late Cretaceous Period. It is part of the Nodosauridae, a family within Ankylosauria. It is named after the Edmonton Formation (now the Horseshoe Canyon Formation in Canada), the unit of rock where it was found. Description Size and general build ''Edmontonia'' was bulky, broad and tank-like. Its length has been estimated at about 6.6 m (22 ft). In 2010, Gregory S. Paul considered both main ''Edmontonia'' species, ''E. longiceps'' and ''E. rugosidens'', to be equally long at six metres and weigh three tonnes.Paul, G.S., 2010, ''The Princeton Field Guide to Dinosaurs'', Princeton University Press p. 238 ''Edmontonia'' had small, oval ridged bony plates on its back and head and many sharp spikes along its sides. The four largest spikes jutted out from the shoulders on each side, the second of which was split into subspines in ''E. rugosidens'' specimens. Its skull had a pear-like shape when viewed ...
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