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Angolasaurus At The Smithsonian
''Angolasaurus'' ("Angola lizard") is an extinct genus of mosasaur. Definite remains from this genus have been recovered from the Turonian and Coniacian of Angola, and possibly the Coniacian of the United States, the Turonian of Brazil, and the Maastrichtian of Niger. While at one point considered a species of ''Platecarpus'', recent phylogenetic analyses have placed it between the (then) plioplatecarpines ''Ectenosaurus'' and ''Selmasaurus'', maintaining a basal position within the plioplatecarpinae. Its wide geographic range make it the one of the only Turonian mosasaurs with a transatlantic range. Description ''Angolasaurus'' was a small mosasaur, with a total length of about 4 meters (13 feet). It shared much of a body plan with its relative ''Platecarpus'', but with a slightly longer skull relative to body length. Its skull housed 11 maxillary teeth, 4 premaxillary teeth, and 12 dentary teeth. The phylogenetic relationship of ''Angolasaurus'' indicates that individuals of ...
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Turonian
The Turonian is, in the ICS' geologic timescale, the second age in the Late Cretaceous Epoch, or a stage in the Upper Cretaceous Series. It spans the time between 93.9 ± 0.8 Ma and 89.8 ± 1 Ma (million years ago). The Turonian is preceded by the Cenomanian Stage and underlies the Coniacian Stage. At the beginning of the Turonian an oceanic anoxic event (OAE 2) took place, also referred to as the Cenomanian-Turonian boundary event or the "Bonarelli Event". Stratigraphic definition The Turonian (French: ''Turonien'') was defined by the French paleontologist Alcide d'Orbigny (1802–1857) in 1842. Orbigny named it after the French city of Tours in the region of Touraine (department Indre-et-Loire), which is the original type locality. The base of the Turonian Stage is defined as the place where the ammonite species '' Watinoceras devonense'' first appears in the stratigraphic column. The official reference profile (the GSSP) for the base of the Turonian is located in the Roc ...
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Sergipe
Sergipe (), officially State of Sergipe, is a state of Brazil. Located in the Northeast Region along the Atlantic coast of the country, Sergipe is the smallest state in Brazil by geographical area at , larger only than the Federal District. Sergipe borders Bahia to the south and west and Alagoas to the north. Aracaju is the capital and the largest city in the state; the state is divided into 75 municipalities. The state has 1.1% of the Brazilian population and produces only 0.6% of the Brazilian GDP. Geography As with most of the states in northeastern Brazil, inland Sergipe is almost entirely savanna (''caatinga''), and its coastline is characterized by mangroves, swamps and sandy beaches. A small strip of tropical rainforest runs down the coast. The São Francisco River forms its northern boundary, and the drainage of the northern part of the state is northward and eastward to that river. The southern half of the state slopes eastward and is drained directly into the Atlanti ...
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Igdamanosaurus
''Igdamanosaurus'', meaning "lizard from Igdaman", is an extinct genus of Cretaceous marine lizard belonging to the mosasaur family. It is classified as part of the Globidensini tribe (within the Mosasaurinae), and is like the other members of the tribe recognised by its rounded and knob-like teeth. These teeth indicate a highly specialized lifestyle, likely including a durophagous diet. The genus contains a single species, ''Igdamanosaurus aegyptiacus'', from Maastrichtian-aged marine environments of Africa. Its fragmentary fossil remains have been recovered from the Duwi Formation of Egypt, Ouled Abdoun Basin of Morocco, and the Dukamaje Formation of Niger. Description ''Igdamanosaurus'' was a small durophagous mosasaur at a size roughly similar to the closely related globidensin ''Carinodens.'' Its fossils preserve blunt, rounded teeth similar to those of the other members of the Globidensini. These teeth were better suited for crushing armored prey like molluscs and turtl ...
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Globidensini
The Globidensini or Globidentatini are a tribe of mosasaurine mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the tribe, known as "globidensins" or "globidensine mosasaurs", have been recovered from North America, Europe, Africa and Asia. The tribe contains the genera ''Globidens'' (the best studied genus by far), '' Carinodens'', '' Igdamanosaurus'',Schulp, Anne S., et al. "New mosasaur material from the Maastrichtian of Angola, with notes on the phylogeny, distribution and palaeoecology of the genus Prognathodon." On Maastricht Mosasaurs. Publicaties van het Natuurhistorisch Genootschap in Limburg 45.1 (2006): 57-67/ref>Lindgren, Johan. "Dental and vertebral morphology of the enigmatic mosasaur Dollosaurus (Reptilia, Mosasauridae) from the lower Campanian (Upper Cretaceous) of southern Sweden." Bulletin of the Geological Society of Denmark 52.17 (2005): e25/ref> '' Harranasaurus'' and '' Xenodens''. Features of the maxilla and digits make the placeme ...
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Plioplatecarpus
''Plioplatecarpus'' is a genus of mosasaur lizard. Like all mosasaurs, it lived in the late Cretaceous period, about 73-68 million years ago. Description ''Plioplatecarpus'' was a medium-sized mosasaur, measuring long and weighing . The eyes of ''Plioplatecarpus'' are proportionally larger than those of many mosasaur genera, although the skull is relatively short. The larger eyes may be an adaptation to low light conditions like those found in deeper water. It has fewer teeth than most mosasaurs, but they are greatly recurved. This suggests that ''Plioplatecarpus'' would have hunted relatively small prey that it could grab very precisely. The broad distribution of fossil remains in both North America and Europe suggest that it would have been an open ocean predator. Discovery ''Plioplatecarpus'' was first found in Europe by paleontologist Louis Dollo (''P. marshi''), in 1882. It was relatively incomplete, but more fossils would soon turn up. In North America, Edward Drinker Co ...
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Plioplatecarpinae
Plioplatecarpinae is a subfamily of mosasaurs, a diverse group of Late Cretaceous marine squamates. Members of the subfamily are informally and collectively known as "plioplatecarpines" and have been recovered from all continents, though the occurrences in Australia remain questionable. The subfamily includes the genera '' Latoplatecarpus'', ''Platecarpus'', ''Plioplatecarpus'' and '' Plesioplatecarpus''. Plioplatecarpines were small to medium-sized mosasaurs that were comparatively fast and agile compared to mosasaurs of other subfamilies. The first plioplatecarpines appear in the Turonian and are among the oldest of mosasaurs, and the clade persists throughout the Maastrichtian, a period of approximately 24 million years. The subfamily was seemingly heavily affected during a poorly understood middle-Campanian mosasaur extinction event and its genera appear to have faced competition from mosasaurine mosasaurs during the Maastrichtian, leading to a decline in numbers and in diver ...
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Dukamaje Formation
The Dukamaje Formation is a geological formation in Niger whose strata date back to the Late Cretaceous. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampel, David B; et al. (2004). "Dinosaur distribution (Late Cretaceous, Africa)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 604-605. . A wealth of Mosasaur fossils have also been recovered from this formation, particularly from the area around Mt. Igdaman. Fossil content ;Other reptiles * '' Sokotosuchus ianwilsoni'' * '' Palaeophis sp.'' * '' Podocnemis sp.'' * '' Trematochampsa taqueti'' * '' Libycosuchus sp.'' ;Fishes * '' Asteracanthus aegyptiacus''In Touhout (bed 4)
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Angolatitan
''Angolatitan'' (meaning "Angolan giant") is a genus of titanosauriform sauropod dinosaur from the Upper Cretaceous. It is also the first non-avian dinosaur discovered in Angola. The genus contains a single species, ''Angolatitan adamastor'', known from a partial right forelimb. ''Angolatitan'' was a relict form of its time; it was a Late Cretaceous basal titanosauriform, when more derived titanosaurs were far more common. Discovery and naming After the Angolan Civil War ended in 2002, the PaleoAngola project planned the first Angolan palaeontological expeditions since the 1960s. The first of these expeditions started in 2005 to explore Angola's fossil rich upper Cretaceous rocks, leading to the discovery of ''Angolatitan''. The discovery was made by Octávio Mateus on May the 25 near Iembe in the province of Bengo, and excavations were conducted during May and August 2006. ''Angolatitan'' was described by Octávio Mateus and colleagues in 2011. The generic name means "Angolan ...
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Plesiosauria
The Plesiosauria (; Greek: πλησίος, ''plesios'', meaning "near to" and ''sauros'', meaning "lizard") or plesiosaurs are an order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period, possibly in the Rhaetian stage, about 203 million years ago. They became especially common during the Jurassic Period, thriving until their disappearance due to the Cretaceous–Paleogene extinction event at the end of the Cretaceous Period, about 66 million years ago. They had a worldwide oceanic distribution, and some species at least partly inhabited freshwater environments. Plesiosaurs were among the first fossil reptiles discovered. In the beginning of the nineteenth century, scientists realised how distinctive their build was and they were named as a separate order in 1835. The first plesiosaurian genus, the eponymous ''Plesiosaurus'', was named in 1821. Since then, more than a hundred vali ...
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Halisaurinae
The Halisaurinae are a subfamily of mosasaurs, a group of Late Cretaceous marine lizards. They were small to medium-sized, ranging from just under 3 meters in ''Eonatator sternbergi'' to as much as 8 or 9 meters in ''Pluridens serpentis''. They tended to have relatively slender jaws and small, numerous teeth, suggesting a diet of small fish and other prey. Although the skeleton is primitive compared to other Mosasauridae in many respects, halisaurines had the distinctive hypocercal tail of other mosasaurids suggesting good swimming ability, and they persisted alongside other mosasaurs until the end of the Cretaceous. The earliest known remains of halisaurines occur in rocks of Santonian age and the subfamily persists until the latest Maastrichtian. Halisaurines are known from North and South America, Europe, Asia and Africa, indicating a more or less global distribution in the Late Cretaceous. Four genera are currently recognized: ''Eonatator'', ''Halisaurus'', '' Phosphorosaur ...
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Angolachelys
''Angolachelys'' is an extinct genus of African eucryptodiran turtle which existed in Angola during the Turonian stage of the Late Cretaceous. The type species is ''Angolachelys mbaxi''. The type MGUAN-PA includes skull, jaw, and postcranial fragments found in the Tadi Beds of the Itombe Formation. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... after Mateus ''et al.'' (2009). References Cryptodira Prehistoric turtle genera Turonian life Late Cretaceous reptiles of Africa Fossil taxa described in 2009 {{paleo-turtle-stub ...
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Turtle
Turtles are an order of reptiles known as Testudines, characterized by a special shell developed mainly from their ribs. Modern turtles are divided into two major groups, the Pleurodira (side necked turtles) and Cryptodira (hidden necked turtles), which differ in the way the head retracts. There are 360 living and recently extinct species of turtles, including land-dwelling tortoises and freshwater terrapins. They are found on most continents, some islands and, in the case of sea turtles, much of the ocean. Like other amniotes (reptiles, birds, and mammals) they breathe air and do not lay eggs underwater, although many species live in or around water. Turtle shells are made mostly of bone; the upper part is the domed carapace, while the underside is the flatter plastron or belly-plate. Its outer surface is covered in scales made of keratin, the material of hair, horns, and claws. The carapace bones develop from ribs that grow sideways and develop into broad flat plates th ...
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