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Amphibamid
The Amphibamidae are an extinct Family (biology), family of Dissorophoidea, dissorophoid temnospondyls known from Late Carboniferous-Early Permian strata in the United States. Classification Amphibamidae has traditionally included small-bodied, terrestrial dissorophoids. The name is attributed to Moodie (1909), but it was rarely used because it originally referred only to ''Amphibamus''. Similar monogeneric families were also erected for other small, terrestrial dissorophoids (e.g., Doleserpetontidae), and most of the taxa now recognized as amphibamiforms were placed within the Dissorophidae. Clack & Milner (1993) revived the Amphibamidae to include ''Amphibamus, Platyrhinops, Doleserpeton,'' and ''Tersomius'.'' Daly (1994) further expanded the composition of the Amphibamidae to include the newly described ''Eoscopus'' as well as the Early Triassic form ''Micropholis''. She suggested that the micromelerpetids were also amphibamids, which has not been validated by more recent wo ...
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Gerobatrachus
''Gerobatrachus'' is an extinct genus of Amphibamidae, amphibamid temnospondyl (represented by the type species ''Gerobatrachus hottoni'') that lived in the Early Permian, approximately 290 million years ago (Ma), in the area that is now Baylor County, Texas. When it was first described in 2008, ''Gerobatrachus'' was announced to be the closest relative of Batrachia, the group that includes modern frogs and salamanders. It possesses a mixture of characteristics from both groups, including a large frog-like head and a salamander-like tail. These features have led to it being dubbed a frogamander by the press. Some more recent studies place ''Gerobatrachus'' as the closest relative of Lissamphibia, the group that contains all modern amphibians including frogs, salamanders, and caecilians, or place modern amphibians far from ''Gerobatrachus'' within a group called Lepospondyli. Description The only known specimen of ''Gerobatrachus'' is a nearly complete skeleton (USNM 489135) about ...
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Temnospondyl
Temnospondyli (from Greek language, Greek τέμνειν, ''temnein'' 'to cut' and σπόνδυλος, ''spondylos'' 'vertebra') is a diverse order (biology), order of small to giant tetrapods—often considered Labyrinthodontia, primitive amphibians—that flourished worldwide during the Carboniferous, Permian, and Triassic periods. A few species continued into the Jurassic and Cretaceous periods. Fossils have been found on every continent. During about 210 million years of evolutionary history, they adapted to a wide range of habitats, including freshwater, terrestrial, and even coastal marine environments. Their life history is well understood, with fossils known from the larval stage, metamorphosis, and maturity. Most temnospondyls were semiaquatic, although some were almost fully terrestrial, returning to the water only to breed. These temnospondyls were some of the first vertebrates fully adapted to life on land. Although temnospondyls are considered amphibians, many had cha ...
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Pasawioops
''Pasawioops'' is an extinct genus of early Permian dissorophoid temnospondyl within the clade Amphibamiformes. History of study ''Pasawioops'' was first described from the early Permian fissure fills near Richards Spur in Comanche County, Oklahoma. It is known from three skulls from this locality. The holotype is a complete skull (reposited at the Sam Noble Oklahoma Museum of Natural History MNH, while the referred specimens are larger but more incomplete partial skulls (one at the OMNH and the other at the Field Museum). The taxon is also known from one specimen from the early Permian of Texas (reposited at the Museum of Comparative Zoology), being previously misidentified as a specimen of ''Tersomius texensis''. Anatomy ''Pasawioops'' has a relatively long and slender skull for an amphibamiform, with the posterior skull table not abbreviated as in more derived taxa. A tooth-bearing crest is found along the cultriform process, and the pterygoid bears two ridges of teeth ...
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Amphibamiformes
Amphibamiformes is an unranked clade with Dissorophoidea created by Schoch (2018). It encompasses all of the taxa traditionally considered to be "amphibamids" (subsequently restricted to '' Doleserpeton annectens'' and '' Amphibamus grandiceps'' by Schoch), branchiosaurids, and hypothetically lissamphibians under the traditional temnospondyl hypothesis of lissamphibian origins. These taxa are typically small-bodied dissorophoids and form the sister group to Olsoniformes, which comprises dissorophids and trematopids. Diagnosis (1) Palatine and ectopterygoid reduced to narrow struts; (2) interpterygoid vacuity greatly expanded laterally; (3) humerus length:waist ratio of 6:10; (4) basioccipital and supraoccipital absent. Definition The most inclusive clade containing ''Amphibamus grandiceps'' but not '' Dissorophus multicinctus''. Phylogeny Simplified phylogeny of Dissorophoidea from Schoch (2018): The phylogeny of Amphibamiformes has historically been relatively unreso ...
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Georgenthalia
''Georgenthalia'' is an extinct genus of dissorophoid temnospondyl from the Lower Permian. It is an amphibamid which lived in what is now the Thuringian Forest of central Germany. It is known from the holotype MNG 11135, a small, complete skull. It was found in the Bromacker locality of the Tambach Formation. It was first named by Jason S. Anderson, Amy C. Henrici, Stuart S. Sumida, Thomas Martens and David S. Berman in 2008 and the type species is ''Georgenthalia clavinasica''. Phylogeny Cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to d ... after Fröbisch and Reisz, 2008: References Amphibamids Dissorophids Permian temnospondyls of Europe Permian Germany Fossil taxa described in 2008 Prehistoric amphibian genera {{Permian-animal-stub ...
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Branchiosauridae
Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe.Schoch, R.R. 2008. The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae. Journal of Systematic Palaeontology. 6(4):409-431. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories.Schoch, R.R. and Frobisch, N.B. 2006. Alternative Pathways in an Extinct Amphibian Clade. Evolution. 60(7):1467-1475 The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the retention of gills. Geological/paleoenvironmental information Branchiosaurids mostly inhabited Permo-Carbo ...
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Tersomius
''Tersomius'' is an extinct genus of dissorophoid temnospondyl within the family Micropholidae. It is known from the early Permian of North America (Oklahoma and Texas). History of study ''Tersomius'' was first named in 1910 by American paleontologist E.C. Case based on a partial skull collected from the Archer City Formation of Texas. The type species is ''T. texensis'', in reference to its geographic provenance. The brief original description was supplemented by a lengthier redescription of the holotype and description of new material by Carroll (1964). At least two of these specimens do not belong to any species of ''Tersomius'' and were reassigned by Maddin et al. (2013), which has led some authors to consider it as a potential "wastebasket taxon." Material referred to ''Tersomius'' cf. ''texensis'' was reported from the early Permian South Grandfield locality in Oklahoma by Daly (1973). A second species of ''Tersomius'', ''T. mosesi,'' was named by Olson (1970) for Amos M ...
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Branchiosauridae
Branchiosauridae is an extinct family of small amphibamiform temnospondyls with external gills and an overall juvenile appearance. The family has been characterized by hundreds of well-preserved specimens from the Permo-Carboniferous of Middle Europe.Schoch, R.R. 2008. The intrarelationships and evolutionary history of the temnospondyl family branchiosauridae. Journal of Systematic Palaeontology. 6(4):409-431. Specimens represent well defined ontogenetic stages and thus the taxon has been described to display paedomorphy (perennibranchiate). However, more recent work has revealed branchiosaurid taxa that display metamorphosing trajectories.Schoch, R.R. and Frobisch, N.B. 2006. Alternative Pathways in an Extinct Amphibian Clade. Evolution. 60(7):1467-1475 The name Branchiosauridae (“Branchio” in Ancient Greek denoting gills and “saurus” meaning lizard) refers to the retention of gills. Geological/paleoenvironmental information Branchiosaurids mostly inhabited Permo-Carbo ...
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Batrachia
The Batrachia are a clade of amphibians that includes frogs and salamanders, but not caecilians nor the extinct allocaudates. The name Batrachia was first used by French zoologist Pierre André Latreille in 1800 to refer to frogs, but has more recently been defined in a phylogenetic sense as a node-based taxon that includes the last common ancestor of frogs and salamanders and all of its descendants. The idea that frogs and salamanders are more closely related to each other than either is to caecilians is strongly supported by morphological and molecular evidence, they are for instance the only vertebrates able to raise and lower their eyes, but an alternative hypothesis exists in which salamanders and caecilians are each other's closest relatives as part of a clade called the Procera, with frogs positioned as the sister taxon of this group. Origins The earliest batrachians are the stem-frogs ''Triadobatrachus'' and ''Czatkobatrachus'' from the Early Triassic, about 250 mil ...
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Amphibamus
''Amphibamus'' is a genus of amphibamid temnospondyl amphibians from the Carboniferous (middle Pennsylvanian) of North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and the Car .... This animal is considered to have been close to the ancestry of modern amphibians. Its length was about 20 cm. Gallery Image:Amphibamus.jpg, ''Amphibamus grandiceps'' skeleton References Amphibamids Dissorophids Carboniferous temnospondyls of North America Taxa named by Edward Drinker Cope Fossil taxa described in 1865 Prehistoric amphibian genera {{Temnospondyli-stub ...
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Micropholis (amphibian)
''Micropholis'' (Greek mikros''' = small and '''pholis''' = scale) is an extinct genus of dissorophoid temnospondyl. Fossils have been found from the ''Lystrosaurus'' Assemblage Zone of the Karoo Basin in South Africa and are dated to the Induan (Early Triassic). Fossils have also been found from the lower Fremouw of Antarctica.''Micropholis'' is the only post-Permian dissorophoid and the only dissorophoid in what is presently the southern hemisphere and what would have been termed Gondwana during the amalgamation of Pangea. History of study ''Micropholis'' was one of the first dissorophoids to be named by English paleontologist Thomas Huxley in 1859 based on a partial skull. ''Micropholis stowii'' (properly ''Micropholis stowi'')'','' the type species, is named for George William Stow, the South African geologist and ethnologist who discovered the specimen and who proposed that it represented some extinct amphibian. English paleontologist Richard Owen later named a new genus a ...
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Dissorophidae
Dissorophidae is an extinct family of medium-sized, temnospondyl amphibians that flourished during the late Carboniferous and early Permian periods. The clade is known almost exclusively from North America. History of study Dissorophidae is a diverse clade that was named in 1902 by George A. Boulenger. Junior synonyms include Otocoelidae, Stegopidae, and Aspidosauridae. Early in the study of dissorophoids when the relationships of different taxa were not well-resolved and most taxa had not been described, Dissorophidae sometimes came to include taxa that are now not regarded as dissorophids and may have excluded earlier described taxa that are now regarded as dissorophids. Amphibamiforms were widely regarded as small-bodied dissorophids, and at one point, Dissorophidae was also suggested to also include Trematopidae. 19th century In 1895, American paleontologist Edward Drinker Cope named ''Dissorophus'' from the early Permian of Texas. This was the first dissorophid to be d ...
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