Agonum Dolens
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Agonum Dolens
''Agonum dolens'' is a species of ground beetle in the Platyninae subfamily. The species is common throughout Eastern Europe and Central Asia. Description Beetle in length is from . The body is bronze or bronze-black, legs are brownish-red. The third interval of elytron have three, rarely four pores. The prothorax is broad, and is transverse. Ecology Living on shores, especially on Warta river of Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous .... References dolens Beetles of Europe Beetles described in 1827 {{Platyninae-stub ...
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Carl Reinhold Sahlberg
Carl Reinhold Sahlberg (January 22, 1779, Eura – October 18, 1860, Yläne) was a Finnish naturalist, primarily an entomologist specializing in beetles. He was the father of entomologist Reinhold Ferdinand Sahlberg (1811-1874), grandfather of entomologist Johan Reinhold Sahlberg (1845-1920), and great grandfather of Uunio Saalas (1882-1969). Biography In 1818 Carl Reinhold Sahlberg succeeded Carl Niclas Hellenius as professor of economy and natural history at Finland’s then only University in Turku (Åbo), the Academy of Åbo. In 1827 the town and the university were destroyed by fire. The remnants of the natural history collections were taken to Helsinki where the University then moved under the name of Imperial Alexander University in Finland (and eventually became University of Helsinki in 1918). Sahlberg replaced the lost collections, played a major role in establishing a new botanical garden in Helsinki, and with his pupils organised a scientific society "Societas p ...
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Ground Beetle
Ground beetles are a large, cosmopolitan distribution, cosmopolitan family (biology), family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal families. They belong to the Adephaga. Members of the family are primarily carnivorous, but some members are phytophagous or omnivorous. Description and ecology Although their body shapes and coloring vary somewhat, most are shiny black or metallic and have ridged wing covers (elytra). The elytra are fused in some species, particularly the large Carabinae, rendering the beetles unable to fly. The species ''Mormolyce phyllodes'' is known as violin beetle due to their peculiarly shaped elytra. All carabids except the quite primitive flanged bombardier beetles (Paussinae) have a groove on their arthropod leg, fore leg tibiae bearing a comb of hairs used for cleaning their antenna (biology), antennae. ...
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Platyninae
Platyninae is a subfamily of ground beetles (family Carabidae). Genera The subfamily Platyninae contains about 250 genera organized into three tribes: ; Tribe Omphreini Ganglbauer, 1891 : '' Omphreus'' Dejean, 1828 ; Tribe Platynini Bonelli, 1810 : '' Abacetodes'' Straneo, 1939 : '' Achaetocephala'' Habu, 1975 : '' Achaetoprothorax'' Habu, 1978 : '' Aepsera'' Chaudoir, 1874 : '' Agelaea'' Gené, 1839 : '' Agonidium'' Jeannel, 1948 : '' Agonobembix'' Jeannel, 1948 : '' Agonoriascus'' Basilewsky, 1985 : '' Agonorites'' Jeannel, 1951 : '' Agonum'' Bonelli, 1810 : '' Altagonum'' Darlington, 1952 : '' Anchomenus'' Bonelli, 1810 : '' Andinocolpodes'' Perrault, 1991 : '' Andrewesius'' Andrewes, 1939 : '' Aparupa'' Andrewes, 1930 : '' Archagonum'' Basilewsky, 1953 : '' Archicolpodes'' J.Schmidt, 2001 : '' Arhytinus'' Bates, 1889 : '' Atranodes'' Jedlicka, 1953 : '' Atranus'' LeConte, 1847 : '' Austroglyptolenus'' Roig-Juñent, 2003 : '' Beckeria'' Jedlicka, 1931 : '' Blackburnia'' Shar ...
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Eastern Europe
Eastern Europe is a subregion of the Europe, European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russia, which spans roughly 40% of the continent's landmass while accounting for approximately 15% of its total population."The Balkans"
, ''Global Perspectives: A Remote Sensing and World Issues Site''. Wheeling Jesuit University/Center for Educational Technologies, 1999–2002.
It represents a significant part of Culture of Europe, European culture; the main socio-cultural characteristics of Eastern Europe have historically been defined by the traditions of Slavs and Greeks, as well as by the influence of Eastern Christianity as it developed through t ...
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Central Asia
Central Asia, also known as Middle Asia, is a subregion, region of Asia that stretches from the Caspian Sea in the west to western China and Mongolia in the east, and from Afghanistan and Iran in the south to Russia in the north. It includes the former Soviet Union, Soviet republics of the Soviet Union, republics of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan, which are colloquially referred to as the "-stans" as the countries all have names ending with the Persian language, Persian suffix "-stan", meaning "land of". The current geographical location of Central Asia was formerly part of the historic region of Turkestan, Turkistan, also known as Turan. In the pre-Islamic and early Islamic eras ( and earlier) Central Asia was inhabited predominantly by Iranian peoples, populated by Eastern Iranian languages, Eastern Iranian-speaking Bactrians, Sogdians, Khwarezmian language, Chorasmians and the semi-nomadic Scythians and Dahae. After expansion by Turkic peop ...
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Elytron
An elytron (; ; , ) is a modified, hardened forewing of beetles (Coleoptera), though a few of the true bugs (Hemiptera) such as the family Schizopteridae are extremely similar; in true bugs, the forewings are called hemelytra (sometimes alternatively spelled as "hemielytra"), and in most species only the basal half is thickened while the apex is membranous, but when they are entirely thickened the condition is referred to as "coleopteroid". An elytron is sometimes also referred to as a shard. Description The elytra primarily serve as protective wing-cases for the hindwings underneath, which are used for flying. To fly, a beetle typically opens the elytra and then extends the hindwings, flying while still holding the elytra open, though many beetles in the families Scarabaeidae and Buprestidae can fly with the elytra closed (e.g., most Cetoniinae; ). In a number of groups, the elytra are reduced to various degrees, (e.g., the beetle families Staphylinidae and Ripiphoridae), o ...
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Prothorax
The prothorax is the foremost of the three segments in the thorax of an insect, and bears the first pair of legs. Its principal sclerites (exoskeletal plates) are the pronotum (dorsal), the prosternum (ventral), and the propleuron (lateral) on each side. The prothorax never bears wings in extant insects (except in some cases of atavism), though some fossil groups possessed wing-like projections. All adult insects possess legs on the prothorax, though in a few groups (e.g., the butterfly family Nymphalidae) the forelegs are greatly reduced. In many groups of insects, the pronotum is reduced in size, but in a few it is hypertrophied, such as in all beetles (Coleoptera). In most treehoppers (family Membracidae, order Hemiptera), the pronotum is expanded into often fantastic shapes that enhance their camouflage or mimicry. Similarly, in the Tetrigidae, the pronotum is extended backward to cover the flight wings, supplanting the function of the tegmina. See also *Glossary of entomolo ...
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Shore
A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past and present, while the beach is at the edge of the shore, representing the intertidal zone where there is one. In contrast to a coast, a shore can border any body of water, while the coast must border an ocean or a sea. Therefore, in that sense, a coast is a type of shore. However, the word "coast" often refers to an area far wider than the shore, often stretching miles into the interior. Shores are influenced by the topography of the surrounding landscape, as well as by water induced erosion, such as waves. The geological composition of rock and soil dictates the type of shore which is created. Rivieras ''Riviera'' is an Italian word for "shoreline", ultimately derived from Latin ''ripa'' ("riverbank"). It came to be applied as a proper n ...
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Warta
The river Warta ( , ; german: Warthe ; la, Varta) rises in central Poland and meanders greatly north-west to flow into the Oder, against the German border. About long, it is Poland's second-longest river within its borders after the Vistula, and third-longest including the Oder, that flows also across Czech Republic and Germany.Statistical Yearbook of the Republic of Poland 2017
, p. 85-86
Its

Poland
Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populous member state of the European Union. Warsaw is the nation's capital and largest metropolis. Other major cities include Kraków, Wrocław, Łódź, Poznań, Gdańsk, and Szczecin. Poland has a temperate transitional climate and its territory traverses the Central European Plain, extending from Baltic Sea in the north to Sudeten and Carpathian Mountains in the south. The longest Polish river is the Vistula, and Poland's highest point is Mount Rysy, situated in the Tatra mountain range of the Carpathians. The country is bordered by Lithuania and Russia to the northeast, Belarus and Ukraine to the east, Slovakia and the Czech Republic to the south, and Germany to the west. It also shares maritime boundaries with Denmark and Sweden. ...
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Agonum
''Agonum'' is a large genus of ground beetles in the subfamily Harpalinae, tribe Platynini. They are mid-sized to smallish beetles, typically with dark metallic hues – often reddish or bronze, but sometimes black, green etc. The genus is generally native to the Holarctic and the Mediterranean region; their southern limit in Central Asia and the Himalaya region is less well understood, and they seem to range outward a bit out of their core regions (e.g. into East Africa). They are wet-loving throughout their life cycle; for example, the genus is well represented in Ireland, where they are more plentiful than anywhere else in Europe. Subgenera and selected species While some subgenera are accepted by most authors (e.g. ''Europhilus, Olisares'' and of course ''Agonum''), others are of disputed validity. Moreover, subgeneric separation has historically been mostly focussed on European and Japanese species. The following list does not incorporate the most recent reviews and m ...
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Beetles Of Europe
Beetles are insects that form the order Coleoptera (), in the superorder Endopterygota. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 400,000 described species, is the largest of all orders, constituting almost 40% of described insects and 25% of all known animal life-forms; new species are discovered frequently, with estimates suggesting that there are between 0.9 and 2.1 million total species. Found in almost every habitat except the sea and the polar regions, they interact with their ecosystems in several ways: beetles often feed on plants and fungi, break down animal and plant debris, and eat other invertebrates. Some species are serious agricultural pests, such as the Colorado potato beetle, while others such as Coccinellidae (ladybirds or ladybugs) eat aphids, scale insects, thrips, and other plant-sucking insects that damage crops. Beetles typically have a particularly hard exos ...
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