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Achillesaurus
''Achillesaurus'' is a genus of alvarezsaurid theropod dinosaur from the Santonian-age Upper Cretaceous Bajo de la Carpa Formation of Rio Negro, Argentina. It was named in reference to Achilles' heel, because diagnostic features are found there for these animals. The genus was a relatively large, basal alvarezsaurid, and a contemporary of ''Alvarezsaurus''. ''Achillesaurus'' is based on MACN-PV-RN 1116, a partial skeleton including a sacral vertebra, four tail vertebrae, part of the left thighbone, shin and foot, and the left ilium. Agustín Martinelli and Ezequiel Vera, who described the specimen, performed a phylogenetic analysis and found their new genus to be an alvarezsaurid with an unresolved relationship to ''Alvarezsaurus'' and more derived alvarezsaurids. Makovicky, Apesteguía & Gianechini (2012) argued that ''Achillesaurus'' might actually be a junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In bo ...
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Alvarezsaurid
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests. ''Alvarezsaurus'', the type genus of the family, was named for the historian Gregorio Álvarez. History of study Bonaparte (1991) described the first alvarezsaurid, ''Alvarezsaurus calvoi'', from an incomplete skeleton found in Patagonia, Argentina. Bonaparte also named a family, Alvarezsauridae, to contain it. He a ...
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Alvarezsaurids
Alvarezsauridae is a family of small, long-legged dinosaurs. Although originally thought to represent the earliest known flightless birds, they are now thought to be an early diverging branch of maniraptoran theropods. Alvarezsaurids were highly specialized. They had tiny but stout forelimbs, with compact, bird-like hands. Their skeletons suggest that they had massive breast and arm muscles, possibly adapted for digging or tearing. They had long, tube-shaped snouts filled with tiny teeth. They have been interpreted as myrmecophagous, adapted to prey on colonial insects such as termites, with the short arms acting as effective digging instruments to break into nests. ''Alvarezsaurus'', the type genus of the family, was named for the historian Gregorio Álvarez. History of study Bonaparte (1991) described the first alvarezsaurid, ''Alvarezsaurus calvoi'', from an incomplete skeleton found in Patagonia, Argentina. Bonaparte also named a family, Alvarezsauridae, to contain it. He a ...
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Bajo De La Carpa Formation
The Bajo de la Carpa Formation is a geologic formation of the Neuquén Basin that crops out in northern Patagonia, in the provinces of Río Negro and Neuquén, Argentina. It is the oldest of two formations belonging to the Río Colorado Subgroup within the Neuquén Group. Formerly, that subgroup was treated as a formation, and the Bajo de la Carpa Formation was known as the Bajo de la Carpa Member.Sánchez ''et al.'', 2006 At its base, this formation conformably overlies the Plottier Formation of the older Río Neuquén Subgroup, and it is in turn overlain by the Anacleto Formation, the youngest and uppermost formation of the Neuquén Group. The Bajo de la Carpa Formation can reach in thickness in some locations, and consists mainly of sandstones of various colors, all of fluvial origin, with thin layers of mudstone and siltstone in between. Geological features such as geodes, chemical nodules, impressions of raindrops, and paleosols (fossil soils) are commonly found in this ...
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Alvarezsaurus
''Alvarezsaurus'' (; "Alvarez's lizard") is a genus of alvarezsaurid dinosaur from the Late Cretaceous, living in Argentina approximately 86 - 83 million years ago. It was a small dinosaur, measuring long and weighing approximately . It was found in the Bajo de la Carpa Formation and was named by paleontologist José Bonaparte in 1991 after the historian Don Gregorio Alvarez. The type species is ''A. calvoi''. Description ''Alvarezsaurus'' was a bipedal theropod. Like other lightweight theropods, it had a long tail, and its leg structure suggests that it was a fast runner. The most proximal elements of ''Alvarezsaurus'' caudal vertebrae exhibited ventrally sharp centra and the transerve processes of these vertebrae were sub-triangular and laterodistally directed, features seen in other alvarezsaurids like ''Shuvuuia''. Spinal processes were entirely absent or poorly developed, and each caudal vertebra supported short prezygapophyses. The scapula was visibly curved and proporti ...
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Santonian
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The Santonian is preceded by the Coniacian and is followed by the Campanian.Gradstein ''et al.'' (2004) Stratigraphic definition The Santonian Stage was established by French geologist Henri Coquand in 1857. It is named after the city of Saintes in the region of Saintonge, where the original type locality is located. The base of the Santonian Stage is defined by the appearance of the inoceramid bivalve ''Cladoceramus undulatoplicatus''. The GSSP (official reference profile) for the base of the Santonian Stage is located near Olazagutia, Spain; it was ratified by the Subcommission on Cretaceous Stratigraphy in 2012. The Santonian's top (the base of the Campanian Stage) is informally marked by the extinction of the crinoid '' Marsupites tes ...
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Vertebra
The spinal column, a defining synapomorphy shared by nearly all vertebrates,Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa. The basic configuration of a vertebra varies, but the bone is its ''body'', with the central part of the body constituting the ''centrum''. The upper (closer to) and lower (further from), respectively, the cranium and its central nervous system surfaces of the vertebra body support attachment to the intervertebral discs. The posterior part of a vertebra forms a vertebral arch ...
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Cretaceous Argentina
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of the entire Phanerozoic. The name is derived from the Latin ''creta'', "chalk", which is abundant in the latter half of the period. It is usually abbreviated K, for its German translation ''Kreide''. The Cretaceous was a period with a relatively warm climate, resulting in high eustatic sea levels that created numerous shallow inland seas. These oceans and seas were populated with now-extinct marine reptiles, ammonites, and rudists, while dinosaurs continued to dominate on land. The world was ice free, and forests extended to the poles. During this time, new groups of mammals and birds appeared. During the Early Cretaceous, flowering plants appeared and began to rapidly diversify, becoming the dominant group of plants across the Earth by the ...
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Santonian Life
The Santonian is an age in the geologic timescale or a chronostratigraphic stage. It is a subdivision of the Late Cretaceous Epoch or Upper Cretaceous Series. It spans the time between 86.3 ± 0.7 mya (million years ago) and 83.6 ± 0.7 mya. The Santonian is preceded by the Coniacian and is followed by the Campanian.Gradstein ''et al.'' (2004) Stratigraphic definition The Santonian Stage was established by French geologist Henri Coquand in 1857. It is named after the city of Saintes in the region of Saintonge, where the original type locality is located. The base of the Santonian Stage is defined by the appearance of the inoceramid bivalve ''Cladoceramus undulatoplicatus''. The GSSP (official reference profile) for the base of the Santonian Stage is located near Olazagutia, Spain; it was ratified by the Subcommission on Cretaceous Stratigraphy in 2012. The Santonian's top (the base of the Campanian Stage) is informally marked by the extinction of the crinoid '' Marsupites tes ...
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Late Cretaceous Dinosaurs Of South America
Late may refer to: * LATE, an acronym which could stand for: ** Limbic-predominant age-related TDP-43 encephalopathy, a proposed form of dementia ** Local-authority trading enterprise, a New Zealand business law ** Local average treatment effect, a concept in econometrics Music * ''Late'' (album), a 2000 album by The 77s * Late!, a pseudonym used by Dave Grohl on his ''Pocketwatch'' album * Late (rapper), an underground rapper from Wolverhampton * "Late" (song), a song by Blue Angel * "Late", a song by Kanye West from ''Late Registration'' Other * Late (Tonga), an uninhabited volcanic island southwest of Vavau in the kingdom of Tonga * "Late" (''The Handmaid's Tale''), a television episode * LaTe, Oy Laivateollisuus Ab, a defunct shipbuilding company * Late may refer to a person who is Dead See also * * * ''Lates'', a genus of fish in the lates perch family * Later (other) * Tardiness * Tardiness (scheduling) In scheduling, tardiness is a measure of a delay in exe ...
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Synonym (taxonomy)
The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linnaeus was the first to give a scientific name (under the currently used system of scientific nomenclature) to the Norway spruce, which he called ''Pinus abies''. This name is no longer in use, so it is now a synonym of the current scientific name, ''Picea abies''. * In zoology, moving a species from one genus to another results in a different binomen, but the name is considered an alternative combination rather than a synonym. The concept of synonymy in zoology is reserved for two names at the same rank that refers to a taxon at that rank - for example, the name ''Papilio prorsa'' Linnaeus, 1758 is a junior synonym of ''Papilio levana'' Linnaeus, 1758, being names for different seasonal forms of the species now referred to as ''Araschnia le ...
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Synapomorphy
In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to have evolved in their most recent common ancestor. ) In cladistics, synapomorphy implies homology. Examples of apomorphy are the presence of erect gait, fur, the evolution of three middle ear bones, and mammary glands in mammals but not in other vertebrate animals such as amphibians or reptiles, which have retained their ancestral traits of a sprawling gait and lack of fur. Thus, these derived traits are also synapomorphies of mammals in general as they are not shared by other vertebrate animals. Etymology The word —coined by German entomologist Willi Hennig—is derived from the Ancient Greek words (''sún''), meaning "with, together"; (''apó''), meaning "away from"; and (''morphḗ''), meaning "shape, form". Clade analysis T ...
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Phylogenetics
In biology, phylogenetics (; from Greek language, Greek wikt:φυλή, φυλή/wikt:φῦλον, φῦλον [] "tribe, clan, race", and wikt:γενετικός, γενετικός [] "origin, source, birth") is the study of the evolutionary history and relationships among or within groups of organisms. These relationships are determined by Computational phylogenetics, phylogenetic inference methods that focus on observed heritable traits, such as DNA sequences, Protein, protein Amino acid, amino acid sequences, or Morphology (biology), morphology. The result of such an analysis is a phylogenetic tree—a diagram containing a hypothesis of relationships that reflects the evolutionary history of a group of organisms. The tips of a phylogenetic tree can be living taxa or fossils, and represent the "end" or the present time in an evolutionary lineage. A phylogenetic diagram can be rooted or unrooted. A rooted tree diagram indicates the hypothetical common ancestor of the tree. An un ...
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