Achelata
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Achelata
The Achelata is an infra-order of the decapod crustaceans, holding the spiny lobsters, slipper lobsters and their fossil relatives. Description The name "Achelata" derives from the fact that all the members of this group lack the chelae (claws) that are found on almost all other decapods (from the Ancient Greek , = "not", , = "claw"). They are further united by the great enlargement of the second antennae, by the special "phyllosoma" form of the larva, and by a number of other characters. Classification and fossil record The infraorder Achelata belongs to the group Reptantia, which consists of the walking/crawling decapods (lobsters and crabs). The cladogram below shows Achelata's placement within the larger order Decapoda, from analysis by Wolfe ''et al.'', 2019. Achelata contains the spiny lobsters (Palinuridae), the slipper lobsters (Scyllaridae) and the furry lobsters (Synaxidae, now usually included in Palinuridae), as well as two extinct families, Cancrinidae a ...
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Phyllosoma
The phyllosoma is the larval stage of spiny, slipper and coral lobsters (Palinuridae, Scyllaridae and Synaxidae), and represents one of the most significant characteristics that unify them into the taxon Achelata. Its body is remarkably thin, flat, and transparent, with long legs. The phyllosoma larva of spiny lobsters has a long planktonic life before metamorphosing into the puerulus stage, which is the transitional stage from planktonic to a benthic existence. Despite the importance of larval survival to predict recruitment, not much is known about the biology of phyllosoma larvae. In many cases even natural diet is still unclear. Jeffs A. (2007)
Revealing the natural diet of the phyllosoma larvae of spiny lobster. Bull. Fish. Res. Agen. No. 20, 9-13 (review)
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Spiny Lobster
Spiny lobsters, also known as langustas, langouste, or rock lobsters, are a family (Palinuridae) of about 60 species of achelate crustaceans, in the Decapoda Reptantia. Spiny lobsters are also, especially in Australia, New Zealand, Ireland, South Africa, and The Bahamas, called crayfish, sea crayfish, or crawfish ("kreef" in South Africa), terms which elsewhere are reserved for freshwater crayfish. Classification The furry lobsters (''e.g.'' ''Palinurellus'') were previously separated into a family of their own, the Synaxidae, but are usually considered members of the Palinuridae. The slipper lobsters (Scyllaridae) are their next-closest relatives, and these two or three families make up the Achelata. Genera of spiny lobsters include ''Palinurus'' and a number of anagrams thereof: ''Panulirus'', ''Linuparus'', ''etc.'' (Palinurus was a helmsman in Virgil's ''Æneid''.) In total, 12 extant genera are recognised, containing around 60 living species: *''Jasus'' Parker, 1883 * ...
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Palinuridae
Spiny lobsters, also known as langustas, langouste, or rock lobsters, are a family (Palinuridae) of about 60 species of achelate crustaceans, in the Decapoda Reptantia. Spiny lobsters are also, especially in Australia, New Zealand, Ireland, South Africa, and The Bahamas, called crayfish, sea crayfish, or crawfish ("kreef" in South Africa), terms which elsewhere are reserved for freshwater crayfish. Classification The furry lobsters (''e.g.'' ''Palinurellus'') were previously separated into a family of their own, the Synaxidae, but are usually considered members of the Palinuridae. The slipper lobsters (Scyllaridae) are their next-closest relatives, and these two or three families make up the Achelata. Genera of spiny lobsters include ''Palinurus'' and a number of anagrams thereof: ''Panulirus'', ''Linuparus'', ''etc.'' (Palinurus was a helmsman in Virgil's '' Æneid''.) In total, 12 extant genera are recognised, containing around 60 living species: *''Jasus'' Parker, 1883 *''J ...
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Slipper Lobster
Slipper lobsters are a family (Scyllaridae) of about 90 species of achelate crustaceans, in the Decapoda clade Reptantia, found in all warm oceans and seas. They are not true lobsters, but are more closely related to spiny lobsters and furry lobsters. Slipper lobsters are instantly recognisable by their enlarged antennae, which project forward from the head as wide plates. All the species of slipper lobsters are edible, and some, such as the Moreton Bay bug and the Balmain bug (''Ibacus peronii'') are of commercial importance. Description Slipper lobsters have six segments in their heads and eight segments in the thorax, which are collectively covered in a thick carapace. The six segments of the abdomen each bear a pair of pleopods, while the thoracic appendages are either walking legs or maxillipeds. The head segments bear various mouthparts and two pairs of antennae. The first antennae, or ''antennules'', are held on a long flexible stalk, and are used for sensing the env ...
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Reptantia
Reptantia is a clade of decapod crustaceans named in 1880 which includes lobsters, crabs and many other well-known crustaceans. Classification In older classifications, Reptantia was one of the two sub-orders of Decapoda alongside Natantia, with Reptantia containing the walking forms, and Natantia containing the swimming forms (prawns, shrimp and boxer shrimp). However, in 1963 Martin Burkenroad found Natantia to be paraphyletic and invalid, and instead split Decapoda into the two sub-orders of Dendrobranchiata (prawns) and Pleocyemata. Pleocyemata contains all the members of the Reptantia (including crabs, lobsters, crayfish, and others), as well as the Stenopodidea ("boxer shrimp"), and Caridea (true shrimp). Reptantia remains a valid monophyletic grouping, but is now no longer ranked as a sub-order. Anatomy The name Reptantia means "those that walk", and contains those decapods whose primary mode of locomotion is to walk along a surface using the pereiopods rather than swi ...
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Pleocyemata
Pleocyemata is a suborder of Decapoda, decapod crustaceans, erected by Martin Burkenroad in 1963. Burkenroad's classification replaced the earlier sub-orders of Natantia and Reptantia with the monophyletic groups Dendrobranchiata (prawns) and Pleocyemata. Pleocyemata contains all the members of the Reptantia (including crabs, lobsters, crayfish, and others), as well as the Stenopodidea (which contains the so-called "boxer shrimp" or "barber-pole shrimp"), and Caridea, which contains the true shrimp. Anatomy All members of the Pleocyemata are united by a number of features, the most important of which is that the fertilisation, fertilised eggs are Wikt:incubation, incubated by the female, and remain stuck to the pleopods (swimming legs) until the zoea larvae are ready to hatch. It is this characteristic that gives the group its name. Pleocyemata also possess a Lamella (surface anatomy), lamellar gill structure as opposed to the branches found in the Dendrobranchiata. Systematics ...
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Synaxidae
Furry lobsters (sometimes called coral lobsters) are small decapod crustaceans, closely related to the slipper lobsters and spiny lobsters. The antennae are not as enlarged as in spiny and slipper lobsters, and the body is covered in short hairs, hence the name furry lobster. Although previously considered a family in their own right (Synaxidae Spence Bate, 1881), the furry lobsters were subsumed into the family Palinuridae in 1990,. Subsequent molecular phylogenetics studies have confirmed that the furry lobsters genera don't form a natural group and were both nested among the spiny lobster genera in family Palinuridae. The family now includes the two furry lobster genera and ten spiny lobster genera. Taxonomy There are two genera, with three species between them: *''Palinurellus gundlachi'' Von Martens, 1878 – Caribbean furry lobster, found in the Caribbean Sea and the Atlantic coast of South America; named for Juan Gundlach *''Palinurellus wieneckii'' ( De Man, 1881) – ...
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Scyllaridae
Slipper lobsters are a family (Scyllaridae) of about 90 species of achelate crustaceans, in the Decapoda clade Reptantia, found in all warm oceans and seas. They are not true lobsters, but are more closely related to spiny lobsters and furry lobsters. Slipper lobsters are instantly recognisable by their enlarged antennae, which project forward from the head as wide plates. All the species of slipper lobsters are edible, and some, such as the Moreton Bay bug and the Balmain bug (''Ibacus peronii'') are of commercial importance. Description Slipper lobsters have six segments in their heads and eight segments in the thorax, which are collectively covered in a thick carapace. The six segments of the abdomen each bear a pair of pleopods, while the thoracic appendages are either walking legs or maxillipeds. The head segments bear various mouthparts and two pairs of antennae. The first antennae, or ''antennules'', are held on a long flexible stalk, and are used for sensing the env ...
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Cancrinidae
''Cancrinos'' is a genus of fossil crustacean closely allied with the slipper lobsters. One species is known, ''C. claviger'' from the Jurassic of southern Germany. Taxonomy Fossils of ''Cancrinos'' are rare, and their state of preservation is often imperfect. Count Georg zu Münster first described ''Cancrinos'' in 1839, based on material from the Upper Jurassic Solnhofen limestones of southern Germany. He described two species, ''Cancrinos claviger'' and ''C. latipes'', differentiated by the size of the second antennae, but the two are now considered to be synonyms. Further specimens have been discovered in Upper Cretaceous lithographic limestones of Lebanon, and described as a new species, ''C. libanensis''; however, Haug ''et al.'' (2016) made it the type species of a separate genus '' Paracancrinos''. Classification Although Münster was unable to discern any living relatives of ''Cancrinos'' during his original description, Reinhard Förster proposed in 1984 that ''Can ...
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Decapoda
The Decapoda or decapods (literally "ten-footed") are an order of crustaceans within the class Malacostraca, including many familiar groups, such as crabs, lobsters, crayfish, shrimp and prawns. Most decapods are scavengers. The order is estimated to contain nearly 15,000 species in around 2,700 genera, with around 3,300 fossil species. Nearly half of these species are crabs, with the shrimp (about 3,000 species) and Anomura including hermit crabs, porcelain crabs, squat lobsters (about 2500 species) making up the bulk of the remainder. The earliest fossil decapod is the Devonian ''Palaeopalaemon''. Anatomy Decapods can have as many as 38 appendages, arranged in one pair per body segment. As the name Decapoda (from the Greek , ', "ten", and , '' -pod'', "foot") implies, ten of these appendages are considered legs. They are the pereiopods, found on the last five thoracic segments. In many decapods, one pair of these "legs" has enlarged pincers, called chelae, with the legs be ...
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Polychelida
Polychelida is an infraorder of decapod crustaceans. Fossil representatives are known dating from as far back as the Upper Triassic. A total of 38 extant species, all in the family Polychelidae, and 55 fossil species have been described. History Polychelida had traditionally been included in the infraorder Palinura, alongside the spiny lobsters and slipper lobsters (now in the infraorder Achelata). In 1995, Gerhard Scholtz and Stefan Richter of the carried out a phylogenetic study of the "Reptantia", and concluded that "Palinura" was paraphyletic. They therefore abandoned that taxon and introduced instead the new clade Polychelida. Classification Polychelida belongs to the group Reptantia, which consists of the walking/crawling decapods (lobsters and crabs). Polychelida is the sister clade to the infraorder Astacidea, which contains the "true" lobsters and crayfish. The cladogram below shows Polychelida's placement within the larger order Decapoda, from analysis by Wolfe '' ...
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Tricarinidae
''Tricarina'' is an extinct genus of crustaceans in order Isopoda, known from a single incomplete fossil specimen from the Cretaceous of western Iran. It has a flattened body with three longitudinal ridges, which give it its name. Sources The single known specimen of ''Tricarina'' was discovered in a well core that had been bored at a site on the Khuzestan Plain in south-western Iran, at , and at a depth of . The rocks that contain it are calcareous shales, which form part of the Gadvan Formation, and are part of the main sequence of rocks for the production oil and gas around the Persian Gulf. Examination of the foraminiferan fossils show that the shales are Barremian to Aptian in age. Description The fossil of ''Tricarina gadvanensis'' is known from a part and counterpart from a core drilled to make an oil well. It has been deposited in the Carnegie Museum of Natural History, Pittsburgh, Pennsylvania, as specimen number CM 54197. A segment of the cylindrical core was cu ...
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