Branchial arches

Branchial arches or gill arches are a series of paired bony/cartilaginous "loops" behind the throat (pharyngeal cavity) of fish, which support the fish gills. As chordates, all vertebrateembryos develop pharyngeal arches, though the eventual fate of these arches varies between taxa. In all jawed fish (gnathostomes), the first arch pair (mandibular arches) develops into the jaw, the second gill arches (the hyoid arches) develop into the hyomandibular complex (which supports the back of the jaw and the front of the gill series), and the remaining posterior arches (simply called branchial arches) support the gills. In tetrapods, a mostly terrestrialclade evolved from lobe-finned fish, many pharyngeal arch elements are lost, including the gill arches. In amphibians and reptiles, only the oral jaws and a hyoid apparatus remains, and in mammals and birds the hyoid is simplified further to support the tongue and floor of the mouth. In mammals, the first and second pharyngeal arches also give rise to the auditory ossicles.